22 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 97 



pump wall itself is well supplied with muscles. Ar chips offers nothing 

 unusual in either pump or proboscis. 



Excellent development of the sucking pump may be found in such 

 pyralids as Desmia funeralis and Nomophila noctuella. The cibarial 

 dilators are strong and well spaced, although the pharyngeal dilators 

 are limited to a single pair. In other pyralids, such as the common 

 wax moth, Galleria mellonella, and the lesser wax moth, Achroia 

 grisella, the pump is relatively weak, especially in the last-named 

 species. Other species of pyralids were studied, but nothing unusual 

 was found. 



S phingidae : Snodgrass (1935) has described the sucking pump of 

 a Sphinx moth. In Hemaris thy she the structure of the pump is 

 typical of this family (fig. 10 D). The pair of pharyngeal dilators is 

 large and set close together, and with the cibarial dilators, provide the 

 pump with powerful suction. In one species, Smerinthns geniinatus 

 (fig. 10 C), this development of the sucking pump has reached such 

 a point that little space is left for the brain and the suboesophageal 

 ganglion. The proboscis extensor musculature is also reduced to a 

 single pair of extensors. In fact, the anterior arms of the tentorium 

 are curved laterally in order to accommodate the expanded pump. In 

 Darapsa pholiis the pump is of more moderate proportions, although 

 quite well developed. In this species there are two pairs of pharyngeal 

 dilators. A mouth valve or oral valve is common in this family. The 

 dorsal muscles of the salivarium are also easily found in the sphingidae. 



Geometridae: In this family the sucking pump is generally weak. 

 Figure 5 B shows the head of a typical geometrid, Ennomos suhsig- 

 narius. There are three pairs of pharyngeal dilators and three pairs 

 of cibarial dilators, but all are relatively thin muscles. The dorsal 

 salivarium muscles are also evident, although very small. Caberodes 

 confusaris shows about the same pump as Ennomos. The sucking 

 pump of both the males and the females of the spring cankerworm 

 moth, Paleacrita vernata, is very weak, although provided with four 

 pairs of dilators. 



The sucking pump of Haetnatopis grataria, in addition to possessing 

 a pair of hypopharyngeal retractors (fig. 5 A; rhphy), is of interest 

 because of its unusual formation. There are three pairs of pharyngeal 

 dilators, the posterior pair originating posterior to the antennae and 

 passing between the antenna! nerves (fig. 9E). There are also two 

 pairs of cibarial dilators. 



Noctiiidae: Members of this family possess well-developed sucking 

 pumps, of which that of Hcliothis ohsoleta (fig. 11 A) is typical. 

 Laterally, the pump is provided with a sheet of fibers on each side, the 



