DISTRIBUTION OF LYMPHATICS IN SCORP/ENICIITHYS 65 

 5. FACIAL LYMPHATICS. 



As in the trunk region there is a distinct superficial and pro- 

 fundus system. Strange to say Vogt (1, p. 137) is the only 

 anatomist to have definitely described lymphatics arising from 

 the facial region of Pisces. The first canal emptying into the 

 cephalic sinus in the salmon is said to originate on the temporal 

 (pterotic) crest from two trunks coming from the head. The 

 first branch, which is somewhat similar to the vessel described 

 below in Scorpa>nic]ithys as the profundus facial lymphatic 

 trunk, has its source at the anterior angle of the nasal fossa, 

 and passing through the orbit receives branches from the upper 

 part of the face and head. The second branch, which is evi- 

 dently analogous to the superficial facial trunk in Scorpamich- 

 thys, is represented as following along under the suborbital 

 bones and collecting numerous branches from the surface of the 

 cheeks, of which the inferior maxillary vessel is the largest; 

 this is said to run along in front of the preopercle from which 

 it receives several branches. Hyrtl (7, p. 236) describes a 

 swelling of the jugular at the entrance of the optic nerve into 

 the orbit that is in communication with a similar bulb on the 

 opposite side as the sinus ophthalmicus (Fig. 8, d), and this 

 sinus he thinks receives the lymph from the head. In a pre- 

 vious paper (2) a similar sinus-like vessel was described as cross- 

 ing the eye muscle canal and connecting the 2 internal jugular 

 veins ; but with Ophiodon there is no marked swelling of the 

 jugulars at the junction with the connecting vessel, which is 

 evidently nothing more than a venous sinus. Stannius (24, p. 

 254) claims that the connection of the head and trunk lymphatics 

 has not yet been made clear. 



Superficial facial lymphatic trunk (Figs. 4 and 5, S.Fac- 

 L. V.). — With Scorpatnichthys this trunk has its origin in the 

 region of the first suborbital bone from a dorsal and a ventral 

 fork ; the dorsal branch comes from the snout and the space 

 surrounding the nasal sac ; while the ventral branch follows 

 along above and behind the maxilla. After uniting the common 

 stem crosses the orbit between the adductor muscle of the pala- 

 tine arch and the upper and inner edge of the chain of subor- 

 bital bones, or suborbital stay as it is in this species. Upon 



