NEW GENERA AND RACES OF AFRICAN UNGULATES 

 By EDMUND HELLER 



NATURALIST, SMITHSONIAN AFRICAN EXPEDITION 



In attempting to determine the skull-characters of the various 

 species of East African ungulates, several species have been found 

 to possess generic characters different from those of the species with 

 which they have hitherto been associated. They are described in 

 the present paper, together with six new races of antelopes detected 

 among the specimens collected by the Smithsonian African Expedi- 

 tion under direction of Colonel Roosevelt. In making these studies, 

 the material in the British Museum has been drawn upon extensively 

 and has been of invaluable assistance. 



Family EQUID^ 

 DOLICHOHIPPUS, new genus 



Type ; Equus grcvyi Oustalet. 



Characters. — Skull elongate and narrow, the rostal portion and 

 to a lesser degree, the occipital region, produced along the main axis 

 of the skull ; rostral portion long and narrow, the diastema equalling 

 the alveolar length of the four last cheek teeth ; lambdoidal crest of 

 occipitals produced behind condyles and extending almost on a level 

 with the auditory meatus, not vertically above it as in Equus or 

 Hippotigris; facial portion of lachrymal bone large, forming one- 

 third of orbital rim, and extending well behind nasal bones poster- 

 iorly; its length 2 to 2}4 times its greatest depth, interorbital region 

 flat or truncate medially; ascending process of premaxilla narrow; 

 its suture with the nasal lying below the nasal notch which is wholly 

 in the nasal bone ; palatal foramina long and narrow ; brain-case 

 very shallow and depressed ; color pattern composed of numerous 

 narrow black and white transverse stripes, the loins without any 

 "gridiron" pattern; ears large and broad; hoofs large and some- 

 what elongate ; ergots on forelegs usually small. 



The shape of the skull of Dolichohippus grevyi is decidedly 

 dolichocephalic, both the rostral and occipital portions being pro- 

 duced somewhat as in the skull of the white rhinoceros, Ceratothe- 

 rinm. The general shape of the skull is, in fact, much nearer that of 



Smithsonian Miscellaneous Collections, Vol. 60, No. 8. 



