H. WORMALD 191 



On the outer surface of the lobes are glands, which are smaller and 

 less couspicuous than those of the leaves. The number (per each lobe) 

 is by no means constant even on the lobes of the same flower and for 

 any one plant is usually from to 8 per lobe. Occasionally one meets 

 with extreme types hke No. 279 in which the number is frequently 15, 

 a lobe with glands absent altogether being of very rare occurrence, 

 and No. 83 of which the lobes are usually eglandular but may have 

 1 to 3 glands, no more than 3 per perianth lobe having been observed 

 in this plant. In the case of the Oregon males the number is generally 

 well above the average and may reach 20. 



Glands of the anthers. Along the dorsal (outer) furrow of each 

 anther may be found a number of comparatively large glands, arranged 

 in a single row when they are few but frequently biseriate when 

 more than 10 are present. In the English forms the number is from 

 to 8 (usually 2 to 4) ; 9 or 10 are rarely found and 11 were counted 

 on one occasion only. With the exception of the last all anthers with 

 more than 10 glands have been foimd only on the Oregon plants and on 

 seedlings with an Oregon plant as one of the parents. In the Oregon 

 males themselves the number of glands per anther varies from 4 to 18 

 (usually about 10) and the glands are often biseriate in the furrow. 

 Among the English plants none has been conspicuous in producing 

 very few glands except perhaps A 15 and I 31, in each of which the 

 number is usually to 2 and no more than 4 have been seen on an 

 anther of either plant. 



The glands of the disc. The disc from which the stamens arise is also 

 often glandular, but this point was not noticed until too late in the season 

 of 1912 for full observations to be made that yeari though it was given 

 attention during the two succeeding years. These glands are minute 

 and individually invisible to the naked eye, therefore easily overlooked 

 except when numerous. Here again the number often varies con- 

 siderably in the same plant, from to 15 being a range of variation 

 frequently encountered at a hill ; yet extreme types are to be recognized. 

 Thus one plant (C 3) has borne glands from 10 to 25 in number, another 

 (I 17) 5 to 25, and in these they are quite conspicuous when observed 

 by means of a hand-lens; in the Oregon <? hop on the other hand 

 no disc-glands have yet been discovered and apparently are not 

 developed in that plant. This feature of the Oregon male is of interest 

 since in the case of other glandiferous organs, viz. leaves, anthers, 



' Afterwards it was found that these glands are figured in Braungart's Der Hopfeit, 

 p. 200. 



