250 Journal of Agricultural Research voi. xvi. N0.9 



Table IV shows the decided change in form produced by the application 

 of V. planifolia pollen to the vanillon stigma. Without exception the 

 apical girth of the fruits from crossed flowers was greater than the basal 

 girth. The average girth at i inch from blossom end was 28.2 per cent 

 greater than that at i inch from stem end. Not only was this relative 

 difference evident in the development of the two ends of the crossed fruits 

 but while the development of the base of the crosses fell far below that of 

 the close-fertilized fruits, the development of the apex in the former ex- 

 ceeded that of the latter for the fruits measured. 



Plate 32, A, shows V 13 fertilized by V. planifolia in the upper row 

 with close-fertilized fruits of V 1 3 in the lower row. Figure B shows the 

 same fruits with sections cut at the lines of measurement. These sec- 

 tions show the fertilization in the two ends to be much more uniform 

 for the close-fertilized fruits than for the crossed fruits which show a 

 heavier fertilization near the apex than near the base. 



In Plate 33, A and B, at the right are shown four fruits of V 34 from 

 blossoms fertilized by V. planifolia. The four at the left are from close- 

 fertilized blossoms of V 34. The heavy fertilization of ovules in the apex 

 of the crosses is clearly shown. 



In Plate 34, A and B, are shown fruits of V 43, the upper row from 

 blossoms fertilized by V. planifolia pollen, the lower row from close- 

 fertilized blossoms. The effect of the V. planifolia pollen was more pro- 

 nounced on this variety than on any other tested, as none of the seven 

 crossed fruits showed any ovules fertilized in the stem end, in some in- 

 stances none for more than 2 inches from the base, though all showed 

 large numbers of ovules fertilized near the blossom end. All of the close- 

 fertilized fruits examined, however, showed many ovules fertilized near 

 the base, fertilization here being frequently heavier than near the blos- 

 som end. In figure B at the top are shown the middle sections of the two 

 fruits at the left in the upper row. The point to which the ovules are 

 fertilized is clearly shown, the ovules appearing as black dots. At the 

 bottom of figure A the middle sections of the two fruits at the left in the 

 lower row show fertilization throughout. 



Plate 35, A, shows longitudinal sections of typical fruits, being from 

 left to right V 9 xV<f,V9XPcf,P9XVd',andP9 XP<f. 



The proportions of the V. planifolia and vanillon blossoms suggest a 

 possible reason for the difference in location of the ovules fertilized by the 

 two kinds of pollen with the resultant alteration in the form of the fruit. 

 At blossoming the difference in the length of the ovaries is slight, but the 

 vanillon column is much longer than that of V. planifolia, exceeding the 

 length of the latter in some instances by as much as 60 to 70 per cent. 

 Plate 35, B, shows a cleared vanillon column and ovary above with that 

 of V. planifolia below, placed to compare the length of ovary at the right 

 and the length of column at the left. 



