ANNOTATED SYSTEMATIC LIST OF 



MARINE TARDIGRADA OF 



THE WORLD 



The arrangement of the following list is based on the 

 classification suggested by Ramazzotti (1972). Original 

 descriptions of all species are available through references in 

 the bibliography. Where subsequent descriptions are useful, 

 they are cited parenthetically. Finally, the most typical 

 habitat and geographical range is indicated for each species. 



Order HETEROTARDIGRADA 

 Suborder Arthrotardigrada 

 FAMILY HALECHINISCIDAE 



Halechiniscus guiteli Richters 1908. Interstitial, sandy 

 intertidal. English Channel, eastern Atlantic, Mediter- 

 ranean, Black Sea. 



Halechiniscus intermedins Renaud-Mornant 1967b. Inter- 

 stitial, subtidal, coralline sand at 4 m depth. Southern 

 Pacific. 



Halechiniscus perfectus Schulz 1955. Interstitial, sandy 

 intertidal and subtidal at 170 m depth. North Sea, Medi- 

 terranean, southern Pacific, Indian Ocean. 



Halechiniscus remanei Schulz 1955 (McGinty 1969). Inter- 

 stitial, sandy intertidal to subtidal at 150 m depth. Medi- 

 terranean, eastern Atlantic, eastern Pacific. 



Halechiniscus subterraneus RenaudDebyser 1959b. Inter- 

 stitial, intertidal in coralline sand. Western Atlantic. 



Pleocola limnoriae Cantacuzene 1951. Commensal of isopod 

 Limnoria lignorum and sandy intertidal to subtidal at 

 130 m depth. English Channel, western Atlantic. 



Actinarctus doryphorus Schulz 1935 (Grell 1937). Intersti- 

 tial, sandy intertidal to subtidal at 170 m depth. One 

 report of possibly ectocommensal relationship with 

 Echinocyamus pusiUus (Grell 1937). North Sea, English 

 Channel, eastern Atlantic. 



Tetrakentron synaptae Cuenot 1892. Ectocommensal on 

 tentacles of holothurian, Leptosynapta galliennei. Eng- 

 lish Channel. 



Styraconyx haploceros Thulin 1942. Lichens near high 

 tidal line. English Channel. 



Styraconyx paulae Robotti 1971. Shallow subtidal, epizoic 

 on coral. 



Styraconyx sargassi Thulin 1942. Pelagic on algae, 

 especially Sargassum. Western Atlantic, Mediterra- 

 nean, Gulf of Mexico, eastern Pacific. 



Bathyechiniscus tetronyx Steiner 1926. Questionable de- 

 scription based on single observation from mud at 400 m 

 depth in south Atlantic. 



Flararctus antiUensis Van der Land 1968 (Renaud-Mornant 

 1971). Interstitial in coralline sand, 3 m depth. Carib- 

 bean. 



Florarctus heimi Delamare Deboutteville and Renaud- 

 Mornant 1965 (Delamare Deboutteville and Renaud- 

 Mornant 1966: Renaud-Mornant 1967b). Interstitial in 

 coralline sand. Southern Pacific. 



Florarctus salvati Delamare Deboutteville and Renaud- 

 Mornant 1965 (Delamare Deboutteville and Renaud 



Mornant 1966: Renaud Mornant 1967b). Interstitial in 

 coralline sand. Southern Pacific. 



Tanarctus arborspinosus Lindgren 1971. Interstitial, inter- 

 tidal. Western Atlantic. 



Tanarctus tauricus Renaud-Debyser 1959b. Interstitial, 

 intertidal. Western Atlantic. 



FAMILY BATILLIPEDIDAE 



Batillipes acaudatus Pollock 1971. Interstitial, sandy inter- 

 tidal. Northestern Atlantic. 



Batillipes annulatus DeZio 1962. Interstitial, sandy inter- 

 tidal. Mediterranean. 



Batillipes huUacaudatus McGinty and Higgins 1968 (Pollock 

 1970a). Interstitial, sandy intertiday. Northwestern and 

 western Atlantic. 



Batillipes camonensis Fize 1957. Interstitial, sandy inter- 

 tidal. Mediterranean, Indian Ocean. 



Batillipes dicrocercus Pollock 1970a. Interstitial, sandy 

 intertidal. Northwestern Atlantic. 



Batillipes friaufi Kiggin 1962. Interstitial, sandy intertidal. 

 Western Atlantic. 



Batillipes gilmartini McGinty 1969. Interstitial, sandy 

 intertidal. Eastern Pacific. 



Batillipes littoralis Renaud-Debyser 1959a. Sandy inter- 

 tidal, interstitial. Eastern Atlantic. 



Batillipes minis Richters 1909 (Marcus 1927; Rudescu 1964: 

 McGinty and Higgins 1968: Pollock 1970a, b). Sandy in 

 tertidal and shallow subtidal, interstitial. Baltic, North 

 Sea, Black Sea, Mediterranean, Atlantic, Indian Ocean. 



BatiUipes pennaki Marcus 1946 (DeZio 1962: Pollock 1970a, 

 b). Interstitial, sandy intertidal. Northern and south- 

 ern Atlantic, Mediterranean, Indian Ocean. 



5atiMipespArea<tcus Renaud-Debyser 1959a (Riemann 1966: 

 Pollock 1971). Intertidal, interstitial in sand. North- 

 eastern Atlantic, North Sea. 



Batillipes similis Schulz 1955. Sandy intertidal, interstitial. 

 Mediterranean. 



Batillipes tubematis Pollock 1971 (Riemann 1966). Sandy 

 intertidal, shallow subtidal. North Sea, northeastern 

 Atlantic. 



Orzeliscus belopus Bois-Reymond Marcus 1952 (Renaud- 

 Mornant 1967b). Interstitial in sand, intertidal and shal- 

 low subtidal. Northeastern, western, and southwestern 

 Atlantic. 



Orzeliscus septentrionalis Schulz 1953b. Interstitial in 

 sand, intertidal. North Sea. 



FAMILY CORONARCTIDAE 



Coronarctus tenellus Renaud-Mornant 1974. Abyssal mud. 

 Southern Atlantic, Indian Ocean. 



FAMILY STYGARCTIDAE 



Stygarctus bradypus Schulz 1951 (Renaud-Mornant and An- 

 selme-Moizan 1969). Interstitial, intertidal. North Sea, 

 northwestern and northeastern Atlantic, southern Pacific. 



Stygarctus granulatus Pollock 1970a (Pollock 1970b). In- 

 terstitial, intertidal. Northwestern Atlantic. 



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