34 Genetic Studies in rntiifnes: Sferilifif 



luulor baj;s. The seed from tlip natural berries was sowu separately 

 ,i;ivin<^ 150 seedlings; but eight plants from one berry proved from other 

 evidence to be the result of a cross and were therefore not included, 

 otherwise the results are consistent and trustworthy. Twenty-one plants 

 flowered and were classified as follows: 



Abundant ... 8 Small ... ... 3 



Medium ... G Nil I 



The qualitative results are shown in Table I. It was noted that 

 even the fertile plants, unlike those from Edgecote Purple selfed, were 

 apt to vary considerably in different flowers and on different dates both 

 in quantity and quality of pollen. Although itself fertile Edzell Blue 

 selfed actually throws a large proportion of .sterile offspring, a result 

 which contrasts strongly with that obtained from Edgecote Purple. A 

 somewhat similar appearance of sterility is recorded by Salaman(i) in 

 certain lines derived from a fertile Sutton's Flourball. We shall .submit 

 an explanation after the much fuller evidence from crosses has been 

 given. 



Two hundred seedlings were raised from the cross — Edgcote Purple x 

 Myatt's Ashleaf in 1921. Of the F^ plants 22 flowered, consisting of 



Abundant ... 19 Medium ... 3 



All were fertile and contained at least 20 per cent, of good grains. 



A similar number of seedlings were raised from the reciprocal cross 

 — Myatt's Ashleaf ■: Edgcote Purple — of which 13 flowered. All were 

 fully fertile. The results from this cross are at once intelligible and 

 harmonise with the few selfed data available, fertility behaving as a 

 recessive. 



The cross — Edzell Blue x Myatt"s Aslileaf — was raised in 1920, and 

 grown on in 1921 when a further sowing was also made. Of the 163 

 plants grown 25 flowered and were recorded as: 



Abundant jiollen 8 Small ... ... 3 



Medium jjoileu 4 Very few grains 10 



As in the case of Edzell Blue selfed the quantity varied considerably in 

 different records from the .same individual. Unfortunately the quality 

 of pollen here was only examined in three cases (see Table I). It is 

 clear, however, that a large proportion of sterile plants result from this 

 cross between two fertiles. 



In 1919 the crosses — Edzell Bhie x Edgecote Purple and Edgecote 

 Purple ;< Edzell Blue — were made. In the families raised in 1920 there 



