NO. 4 NERVOUS SYSTEM OF A CENTIPEDE — LORENZO 3 



Subclass Epimorpha. 

 Orders : 



Scolopendromorpha. 

 Geophilomorpha. 



Count Carl von Attems (1926), however, makes several modifications. 

 He does not consider the Scutigeromorpha worthy of subclass rank on 

 the basis of its gross anatomical features alone but respects the em- 

 bryological evidence as justifying his division of the order into the 

 subclasses Anamorpha and Epimorpha. He further relegates the 

 Craterostigmophora to subordinal rank. His is the following classifi- 

 cation : 



Subclass Anamorpha. 

 Orders : 



Scutigeromorpha. 

 Lithobiomorpha. 

 Suborder Craterostigmophora. 

 Subclass Epimorpha. 

 Orders : 



Scolopendromorpha. 

 Geophilomorpha. 



The classification of Attems appears to be the more natural and is the 

 scheme which we have adopted for our discussion in the light of the 

 following evidence: 



Anamorpha. — Postembryonic development is by hemianamorpho- 

 sis — that is, the young leaves the egg with only seven fully formed 

 trunk segments and seven pairs of legs ; the eggs are laid singly by the 

 parent female and she broods neither the eggs nor the young. The 

 adults of both orders possess 15 pairs of ambulatory appendages. The 

 Scutigeromorpha is the only order of the class possessing compound 

 eyes, and the external respiratory openings (stomata) are dorsally 

 situated and unpaired. The Lithobiomorpha have simple ocelli, when 

 visual elements are present, and the external respiratory openings 

 (stigmata) are lateral and paired. 



Epimorpha. — Postembryonic development is by epimorphosis — that 

 is, the young animal leaves the egg with the adult complement of pedal 

 segments and pairs of legs ; the eggs are laid in groups, and the female 

 broods both the eggs and the young. The Scolopendromorpha have 

 either 21 or 23 pairs of walking legs, and this number is constant 

 within the species. The antennae have at least 17 articles. The Geo- 

 philomorpha, on the other hand, possess no less than 31 pedal seg- 

 ments and may number as high as 183 with intraspecific variation.* 

 The antennae constantly have 14 articles. 



* It should be remarked that, exclusive of the prehensors, the number of pairs 



