34 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. I40 



Ferris failed to emphasize that Fahlander was describing the nerve 

 as it occurs in the scutigeromorph Thereuopoda. Fahlander made no 

 mention of this nerve's occurrence in the other species he studied — 

 either in the text or in the figures. 



(2) Fahlander described a nerve (ns) which originates between 

 the protocerebrum and deutocerebrum whose destination is the 

 frontal ganglion. Ferris denied its existence : "No such nerve appears 

 in our material. No such nerve should be present. If the generaliza- 

 tions previously offered are valid no such nerve can be present." (P. 

 12.) Fahlander was still, however, referring to the organization of 

 the brain of Thereuopoda. The nerve in question is conspicuously 

 absent in the other three representatives he described and illustrated. 



(3) Fahlander described still another nerve (N17) : 



Die Labralnerven gehen vom Tritocerebrum gleichzeitig mit den Frontalkonnek- 

 tiven ab und bilden einen Plexus praefrontalis ganz kranial in Kopf. Von dort 

 gehen Nerven fiir die clypeale Muskulatur sowie die sensorischen Nerven des 

 Labrum aus. (P. 81.) 



Ferris criticized him principally on the basis of a definition of muscle 

 origin and insertion which first appeared in Applegarth (1952, p. 132). 

 Fahlander made no reference to origin or insertion in the clypeal 

 musculature. It is clear, furthermore, from his own words and from 

 the German idiom, that he was referring primarily to the plexus 

 praefrontalis and not to a direct branch of the labral nerve as Ferris 

 described it. On the contrary, Fahlander wrote, concerning Lithobius: 



Der Frontalnerv (nir,), welcher unpaar ist, aber eine doppelte Wurzel besitzt, 

 verlauft in kranialdorsaler Richtung und innerviert die Clypeusmuskulatur. Er 

 hat keinerlei Verbindimg mit den Labralnerven (ni?), die vom kaudalen Teil 

 der Briicke abgehen und sensorisch sind. Ausser zum Labrum schicken die 

 Labralnerven Aste zur lateralen Partie des Clypeus. (P. 85.) (Italics mine. 

 M.A.L.) 



Fahlander is beyond reproof on this point of Ferris's criticism. 



(4) Fahlander described a free tritocerebral commissure in Litho- 

 bius (p. 86 and fig. 29). It is represented as a dorsal and ventral 

 doublet having connections with two other nerves. Ferris, however, 

 denied that such a commissure existed in his material. He suggested 

 that paired stomodaeal nerves innervating the esophagus may have 

 been mistaken for a free tritocerebral commissure. Fahlander was 

 aware that this structure was a subject of controversy and Ferris ad- 

 mitted that it may be present in some species and not in others as had 

 been shown in the insects. Both authors could be correct since they 

 concerned themselves with different species of lithobiomorphs. 



(5) Fahlander claimed that nerves proceeding from the commis- 



