Material Studied: 1 female (2.14 mm), sample 13 

 (Table 3); 1 female (on slide), sample 14; 1 male (1.60 

 mm), sample 50. 



Description 



Pi Ri with 3 inner setae; P2 Ri2 with 1 outer seta. 



Female.— Length 1.98-2.33 mm. Prosome in dorsal 

 view long, slender ovoid; anterior tapering toward 

 slightly truncate head. In lateral view with uneven, 

 gentle forward slope. Th4 and Th5 separate; Thl-Th4 

 without lateral spinules; Th5 lateral corners 

 protruding about one-third length of genital segment 

 (somewhat more than shown by Sars, 1924, fig. 1). 

 Prosome length slightly more than 4 times urosome. 

 Genital segment as long as wide; protruding ventrally 

 one-third depth of rest of urosome. Caudal ramus 

 length slightly more than width. Caudal setae 

 symmetrical; inner terminal seta omitted in Sars' 

 figure. 



Al unknown; broken in all described specimens. 

 [Sars (1925) stated that Al must be longer than the 

 body, but his figure shows Al broken at segment 21 

 next to distal edge of Thl, so that a complete Al 

 would probably not extend beyond Th4.] 



A2 Rel without setae, as shown by Sars. 



Mx2 without spines on base of lobe-5. 



Mxp about as shown by Sars, except 4 setae on Ri2 

 and only 4 on Ri5. Bl without spine-comb; transverse 

 row of short spinules on distal inner edge. B2 without 

 spine-comb; longitudinal row of stiff long hairs on in- 

 ner edge. 



Pi as shown by Sars, except typical short, thin, out- 

 er seta on B2 about two-thirds distal; row of spinules 

 on distal outer anterior surface of Rel-2; no spines on 

 Re3 surface. 



P2 as shown by Sars, except distal outer anterior 

 spinules on Re2; Rel outer spine forming strongly 

 bent hook; no surface spines on Re3, terminal seta 

 with 30-32 primary teeth. 



P3 (Fig. 85) Ri only shown by Sars. Bl with several 

 outer and inner surface spines; inner margin with 

 hairs. Re2 posterior surface with irregular cluster of 

 spines; Re3 posterior surface with 2 distal clusters of 

 small spines; terminal seta with 31 primary teeth. 



P4 incomplete on specimens studied; Bl with 

 setules, not spinules, in position shown by Sars. 



Male.— Length 1.59-1.76 mm. Prosome in dorsal 

 view robust ovoid, anterior and posterior irregularly 

 rounded; in lateral view with abrupt forward slope. Ce 

 and Thl separate; Ce not expanded dorsolaterally. 

 Prosome about 2Vi length of urosome. Anal segment 

 not reduced, one-half length of caudal ramus. 



Al described by Tanaka (1956), typical left-handed 

 appendage; length unknown. Segment 25 not re- 

 duced. 



PI as figured by Wheeler (1970), except typical out- 



er seta on B2, and Ri lobe with 1-3 terminal points. 

 Rel outer spine short, reaching slightly beyond 

 midlength of Re2. 



P2 Rel outer spine typical, not forming strongly 

 bent hook. 



P4 Bl without inner seta. 



P5 essentially uniramus, left-handed; similar to 

 that on S. antarcticus male except rudimentary left 

 Ri. Left Re2 nearly equaling length of Rel. 



Remarks 



Tanaka (1956) described a new species, S. longipes, 

 for one male taken in a sample with three female S. 

 angusticeps. The similarity of P5 to S. antarcticus 

 male P5, and the 3 inner setae on PI Ri on S. 

 angusticeps female (male otherwise unknown) and S. 

 longipes (sample 50, Table 3) definitely relates S. 

 longipes to the group including S. angusticeps, S. ant- 

 arcticus, S. magnus, and S. validus. Males have been 

 assigned to the latter three species, and S. longipes is 

 considered in the present study to be the male of S. 

 angusticeps. Spinocalanus longipes has also been 

 reported by Grice and Hulsemann (1967), who found 

 female S. angusticeps in adjacent areas. Wheeler 

 (1970) reported one male Spinocalanus sp. Johnson; 

 Wheeler's specimen lacked Re on the longest P5, 

 which he said was the right. Wheeler's figure of P5 

 suggests a rudimentary Ri; the figure of PI essentially 

 agrees with male PI described above, with short Rel 

 outer spine. Wheeler reported female S. angusticeps 

 from the sample which contained this male. 



Distribution 



Records of S. angusticeps are summarized below: 



Pacific Ocean 

 Northwest: Tanaka (1953, 1956), 0-1,000 m. 

 Southwest: Present study, 0-1,000 m. 

 Indian Ocean 

 West: Grice and Hulsemann (1967), 750-2,000 

 m. 

 Atlantic Ocean 



Northeast: Sars (1920; 1924, 1925), 0-2,000 m. 

 East: Grice and Hulsemann (1965), 450-3,000 



m. —Roe (1972b), 660-950 m. 

 Northwest: Grice (1963), 700-1,200 m. —Wheeler 

 (1970), 2,000-4,000 m. 



(14. Spinocalanus caudatus Sars, 1920) 



See Spinocalanus oligospinosus Park, 1970. 



(15. Spinocalanus abyssalis var. 

 pygmaeus Farran, 1926) 



See Spinocalanus longicornis Sars, 1900. 



43 



