Prosome length 2 times urosome. Caudal rami slightly 

 longer than wide, symmetrical. Caudal setae in- 

 complete on specimens examined, bases symmetri- 

 cal. 



Al reaching middle of urosome segment 2; left seg- 

 ment 20 merging with segment 21, typical of right- 

 handed males; segment 25 reduced. IV-2s, le; 

 Vni-2s, 2e; XV-ls, le; XIX-ls, le; right XX-ls; left 

 XX-0; remainder as Table 4. 



A2 Re! with 1 seta. 



Mn blade (Fig. 126) reduced; dorsal tooth relatively 

 long. 



Mx2 long terminal setae much thicker than on 

 female. 



Mxp (Fig. 127) Bl setae of reduced length, surface 

 nude. B2 with longitudinal row of stiff hairs. Ri2-5 in- 

 ner setae inflated about one-third length; Ri2 setae 

 and Ri3 proximal seta distally plumose; other inflated 

 setae nude. Ri2 apparently with only 3 setae, Ri3-5 

 apparently with only 2 setae. Ri4-5 outer setae and 

 Ri5 terminal seta apparently absent. No spinules on 

 inner Ri5. 



PI (Fig. 128) Bl with only few hairs on inner 

 margin. B2 anterior surface with outer distal row of 

 spinules; inner edge with setules and hairs. Re and Ri 

 surface nude. Re3 terminal seta not coarsely ser- 

 rate. 



P2 as on female except Re2 surface spines longer, 

 and Re3 terminal seta with 66-69 primary teeth. 



P3 as on female except Re3 outer spines longer, and 

 terminal seta with ca. 73 primary teeth. 



P4 Bl nude (one specimen with rudimentary right 

 inner seta). Re2-3 missing on specimens examined, 

 remainder as on female. 



P5 (Fig. 129) biramus, right-handed; reaching end 

 of urosome segment 2. Right Bl reaching three- 

 fourths length of left B2; right B2 reaching left Re2. 

 Right leg longer than left; right and left Re, including 

 terminal blade, about equal length; right and left Ri 

 about equal length. Order of length, longest to 

 shortest, of Re segments: left 2, 3, 1; right 1 = 2, 3. 

 Left Ri reaching middle of left Re2. Right Ri reaching 

 one-third length of right Re2. Each Rel with short, 

 flat seta on distal outer edge. Left Re 1-2 more or less 

 fused; inner edge of Rel with few setules. Inner edge of 

 right Re and Ri with hairs. Each Re with 1 small and 1 

 moderate bladelike terminal setae. 



Remarks 



Brodsky (1950) based his description of S. polaris 

 solely on females, and considered S. polaris male as 

 one of two types of male S. elongatus. Brodsky and 

 Nikitin (1955) reported male S'. polaris, so that they 

 may have recognized this error. 



Park's (1970) lengths of females (1.02-1.06 mm) 

 have been questioned in the above description; these 

 should be verified, in view of the differences between 

 some of his published lengths and present 

 measurements of the same specimens. 



Distribution 



Records of S. polaris are summarized below: 



Arctic Ocean 



Central: Brodsky (1950, 1957), 200-3,000 m. 

 Canadian Basin: Brodsky and Nikitin (1955), 



0-3,826 m. —Dunbar and Harding (1968), 



900-3,000 m. — Vidal (1971). —Present studv, 



1,500-3,000 m. 

 Atlantic Ocean 



East: Grice and Hulsemann (1965), 1,500-5,000 



m. 

 Caribbean Sea: Park (1970), 980-2,800 m. 



Brodsky (1957) recorded S. polaris, with some 

 doubt, from the north and northwest Pacific. Some 

 Mimocalanus species look like S. polaris, and if 

 specimens are damaged, they could easily be con- 

 fused. 



27. Spinocalanus brevicaudatus Brodsky, 



1950 



(Figures 130-136, 157) 



Spinocalanus longicornis. — Sars, 1901, p. 22-23, pi. 



12. Not S. longicornis Sars, 1900. 

 Spinocalanus abyssalis. — Sars, 1903, p. 157-158, 



suppl. pi. 3, fig. 2. — van Breemen, 1908 (part), p. 



28-29, fig. 27a, c (only). —With, 1915, p. 69-71, fig. 



15; pi. 1, fig. lOa-b. —Rose, 1933, p. 84-85, fig. 42. 



— Tanaka, 1937 (part), p. 253-254, fig. 4d-e; male 



only. —Davis, 1949, p. 21-22, fig. 23-24. — Farran 



and Vervoort, 1951a (part), p. 2-3, fig. la-d, i (only). 



—Grice and Hulsemann, 1965, p. 229, fig. 7a. Not 



S. abyssalis Giesbrecht, 1888. 

 Spinocalanus major Esterly, 1906, p. 55. 

 Spinocalanus pseudospinipes Brodsky, 1950, p. 127- 



128, fig. 46. 

 Spinocalanus sp.? Brodsky, 1950 (part), p. 130-131, 



fig. 50; male only. 

 Spinocalanus brevicaudatus Brodsky, 1950, p. 134- 



136, fig. 55. — Semenova, 1962, p. 1571-1574, fig. 1- 



4. 

 Spinocalanus similis var. profundalis Brodsky, 1955 



(part), p. 185-187, fig. la-c; female only. 



Type Specimen: Lectotype, female (1.60 mm) OSLO 

 F4923a. 



Type Locality: Northeast Atlantic (Osterfjord), 400- 

 600 m. 



Material Studied: 3 females (1.55, 1.57, 1.60 mm), 

 sample 3 (Table 3); paralectotype, female (1.60 mm, 

 after Sars, 1901), on slide, sample 16; paralectotype, 

 female, on slide, sample 17; lectotype, female (1.60 

 mm) and 1 male stage V (1.40 mm), sample 18; 2 

 paralectotypes, females (1.60 mm), sample 19; 

 paralectotype, male (1.60 mm, after Sars, 1903), on 

 slides, sample 20; 1 female (1.42 mm), sample 21; 1 



53 



