PI (Fig. 220) Rel outer distal edge with 1-3 small 

 teeth. Re2 outer spine not reaching beyond two-thirds 

 length of Re3. Re3 outer spine narrow. Ri outer lobe 

 moderate, with small terminal point. 



P2-P4 (Figs. 221-223) variously incomplete on 

 specimens examined in this study. P4 Bl with inner 

 transverse row of thin, uniform setules. 



Male. — Length of only known specimen 1.28 mm 

 (Tanaka, 1956). According to Tanaka: right P5 with 4 

 segments, but probably Rel-2 more or less fused; Re3 

 relatively large (Tanaka, fig. 13i); left P5 exceeding 

 right by Re3 and one-half length of Re2. 



Remarks 



Tanaka (1956) described two females and one male 

 M. cultrifer; his illustration (fig. 13f) of P2 Ri2 

 suggests an outer seta, not possessed by M. cultrifer. 

 In other respects, also, his description is consistent 

 with that of M. heronae. 



Boucher and de Bovee (1970) reported the first 

 Mimocalanus species from the Mediterranean Sea, 

 four females which were identified as M. distinc- 

 tocephalus. Their description, including figures of 

 complete P2-P4, is consistent with that of M. heronae, 

 except they stated that Th4 and Th5 were separate, a 

 characteristic that may be variable, as in some 

 Spinocalanus species. They also stated that P2 Ri2 

 and P3-P4 Ri3 have 2 outer setae; their figure of P2 

 suggests 1 outer seta, and their figures of P3-P4 in- 

 dicate 1 outer seta, consistent with M. heronae. Their 

 Table 3 indicates 1 outer seta on P2 B2, which should 

 instead be indicated for PI B2. Perhaps the 

 Mediterranean specimens reported by Grice (1971) as 

 M. cultrifer are also M. heronae (see M. cultrifer). 



Distribution 



Minoda (1971) reported two females (1.55, 1.95 

 mm) said to be equivalent to Tanaka's species. These 

 are also considered M. heronae in the present study. 



Vervoort (1946) reported M. cultrifer females up to 

 1.91 mm, from the Indo-Pacific; it is possible that the 

 specimens longer than 1.70 mm are M. heronae. 



Pacific Ocean 



North: Minoda (1971), 743-1,230 m. 



Northeast: Present study, 300 m. 



Central: Present study, 0-1,000 m. 



Northwest: Tanaka (1953, 1956), 300-500 m. 

 —Minoda (1971), 195-485 m. 



Indo-Pacific: ?Vervoort (1946), 636-900 m. 

 Atlantic Ocean 



Mediterranean Sea: Boucher and de Bovee 

 (1970), 50-500 m. 



Etymology: I am pleased to name this species for 

 Gayle A. Heron, Department of Oceanography, 

 University of Washington, Seattle, in gratitude for her 



unfailing friendship over nearly 20 years and in 

 recognition of her enthusiasm and skill in the study of 

 the Copepoda. 



GteNus Teneriforma Grice and 



HULSEMANN,1967 



Tanyrhinus Farran, 1936. 



not Tanyrhinus Mannerheim, 1852. 



Type SPECIES: Teneriforma naso (Farran, 1936). 



Description 



Rostrum a single long blunt cone. PI Rel without 

 outer distal spine; Rel-2 together only slightly longer 

 than Re3. Pi Ri with 2 inner setae. P2 Ri2 with 1 outer 

 seta. P3-P4 Ri3 with 2 outer setae. Surface of swim- 

 ming leg segments without spines. Caudal rami 

 symmetrical, length 2-2' 2 times width. 



Female. — Prosome in dorsal view slender ovoid; 

 head rounded. In lateral view with very abrupt 

 forward slope. Th5 lateral comers prolonged, reaching 

 one-third length of genital segment. Thl-Th4 without 

 lateral spinules. Th4 and Th5 separate. Prosome 

 length 3-3V3 times urosome. Genital segment 

 protruding ventrally one-fourth to one-third depth of 

 rest of urosome. 



Al reaching anal segment; segment 2 length about 

 equaling segment 1; segments 8 and 9 fused. 



A2 Re length about equaling Ri. Rel with 1 seta; 

 Re2 and Re3 fused; Re2 with 2 setae. 



Mn B2 with 3 inner setae. 



Mxp Bl-2 without transverse spine-comb; B2 ap- 

 parently with only 2 midlength setae, and without 

 longitudinal row of stiff hairs or other armament. 

 Published illustrations of Ril-3 indicating less setae 

 than typical for the family. Ri4-5 outer setae 

 moderately long, plumosity unknown. 



P2-P4 Re terminal seta with moderately coarse 

 serrate outer edges; outer flange narrow. 



P4 Bl posterior surface apparently withouf inner 

 transverse row of hairs or setules. Inner seta present. 



Male. — Prosome in dorsal view elongate ovoid; 

 head rounded. In lateral view with very abrupt 

 forward slope. Th5 lateral comers prolonged, as in 

 other male Spinocalanidae; nearly reaching urosome 

 segment 2. Ce not expanded. Ce and Thl separate. 

 Prosome length 3 times urosome. Anal segment not 

 reduced, length two-thirds times caudal ramus. 



Al reaching to end of caudal rami; segment 2 length 

 about equaling segment 1; segments 9 and 10 

 separate; segments beyond 10 not fused; right Al miss- 

 ing from specimen examined; left segments 20 and 21 

 distinct, as in left-handed species; segment 25 not 

 reduced. 



P5 (Fig. 224) uniramus, left-handed, very 

 asymmetrical. Right B2 reduced to small knob; no 



82 



