\E\V Sl'KClES OK CKAWI.INU MKIH'sA KK'OM AllSTHAI.lA l:i;|i;US. 103 



Jii transverse sections tlironfjcli a male of ('. Imsiri'lli at the level of 

 tlie ectodeiinal partitions above tlie radial canals, the testeB are seen to be 

 reduced to inter-radial masses, eaeli oF wliicli is partly cleft by a vei'tical 

 slit into two adradial portions lying- in the aiif^le between the stomach 

 diverticula. Vanh()ft"en describes and Hguies a similar airangement of 

 the testes in tlie Kei'guelen species, and the same condition is shown in 

 Gilelu'ist's figures of transverse sections through a male Medusa. 



In the female of (\ litimi-dH, the ovaries occupy a corresponding 

 position in the subumbrellar cavity to that occupied by the testes in the 

 male. The ovaries, however, show no ti-ace of segregation into two 

 adradial parts. The ova aie closely packed together and, when mature, 

 nieasuie 0-06 mm. in diameter. In the females of the Kerguelen species, 

 Vanluiffen found scarcely any trace of segregation of the ovai'ies into two 

 adradial pai'ts although he suspects that originally the ovaries were 

 separate, and that later, through the growth of the ova, they became 

 merged for lack of space. 



Gilclirist has apparently misintei'preted Vanhoffen's remarks on the 

 ari'angenient of the gonads in ('. kergtieleHeii.'^e. Gilchrist writes : — 

 " Whether or not such partitions between tlie gonads exist in other 

 southern Eleutheria is not known, except in the case of the male of 

 Vanhoffen's species. The females of this species do not appear to liave 

 them, though Vanhoffen suspects they may be present in the young 

 females." Vanhoffen, however, does not meution the presence of partitions 

 or septa between the gonads in the Kerguelen species ; his remarks refer 

 simply to the tendency in the male for the testes to occur as two adradial 

 portions, which are united by a narrow bridge of ectoderm i)i. the angle 

 (" in dem Winkel ") between the stomacli diverticula. Gilchrist lias 

 evidently mistaken this bridge of ectoderm (" Ektodermbriicke ") for a 

 septum ; the true septa, however, are clearly figured b}' Vanhoffen ou 

 Plate XXX., fig. 5c, where they are shown as double folds of ectoderm with 

 a middle one of mesogloea. 



Asexual re^jnx?»c^('oM.— Besides producing gonads, G. ha>!irelli multiplies 

 asexually by budding, the buds arising from the circular canal between the 

 tentacles and the nematocyst ring. Gilchrist found medusa buds occupying 

 a similar position in young specimens of the Cape species. Although the 

 buds were not observed in living specimens of the Medusa from Port 

 Jackson, transvel'se sections of a male revealed six, all of which have reached 

 about the same phase of development. Only very young buds are present 

 and these consist of hollow outgrowths which are nearly cylindrical or 

 slightly tapered at their distal ends. The ectoderm of the bud is composed 

 of several layers of cells. The endodermal cells, in which pigment-granules 

 are recognisable, are arranged in a single iiniforni layer lining the narrow 

 lumen of the bud. 



Affinities. — The Medusa fi'om Port Jackson has been referred to the 

 genus Gnidoneiiia, on account of its crawling habits ; the absence of a brood- 

 pouch above the stomach ; the distinct sexes ; the numerous tentacles, which 

 increase with age and do not correspond in number with the radial canals; 

 the dichotomous tentacles, the upper branch of which carries several 



