162 THE JOURNAL OP BOTANY 



larger than in adnatum, being about 3 mm. long ; it frequently 

 protrudes at the tip of the flower before anthesis. The anthers 

 seem to be usually contabescent. Flowering goes on until very 

 late in the autumn. 



In no case was seed set in 1909-1910, though superficially the 

 ovules appeared well-formed ; and attempts to fertilize the flowers 

 with pollen from adnatum f. stenopliylla and from hirstctum were 

 also fruitless. 



While most of the characters of the hybrid are intermediate 

 between those of the parents, it may be remarked that the pre- 

 sence of long runners, of erect, long, shaggy hairs, of terminal 

 glands on the shorter hairs, and of large flowers (all characters 

 possessed by hirsutum), are all apparently recessive. The study 

 of their exact hereditary behaviour is, however, checked by 

 sterility. 



E. adnatum Griseb., female X montanum Linn., male. The 

 parent stocks originated from plants self-sown in my garden at 

 Tewkesbury, and came true from seed. Both were quite typical, 

 agreeing well with Haussknecht's descriptions. I raised twenty- 

 five plants of the hybrid, and all were absolutely uniform in 

 appearance ; they correspond closely on the whole with Hauss- 

 knecht's descriptions of wild specimens. The following points 

 of interest may be added to his remarks (see Monograph, 

 p. 104) :— 



(1) The upper part of the stem is practically terete, only the 

 lower part showing decurrent lines. 



(2) A transverse commissure joins the bases of the opposite 

 leaves, this being a character derived from montanum. 



(3) The stigma is very shortly four-lobed, this condition differ- 

 ing from that found in adnatum i. stenophylla x hirsutwn (above). 

 Thus the results of crossing § Synstigma and § Schizostigma are 

 not always identical as regards the characters of the stigma of 

 the hybrid. 



(4) The capsules appear well-formed, but the seeds seem to be 

 always imperfect and are without embryos. This ^vas also ob- 

 served by Haussknecht, who states that the seeds are totally 

 sterile. 



The last point is of interest with regard to Bell Salter's artifi- 

 cial hybrid between montanum and " tetragonum " {loc. cit.). He 

 says: " I saved seeds from the original hybrids, and sowed them. 

 The second race was indistinguishable from the first. The seeds 

 of these I again saved and sowed, and still no difference could be 

 detected ; and so on to four turns, when, being satisfied of the 

 reproductive powers of the hybrids and the permanence of the 

 form, I discontinued the experiment." It is difficult to know 

 what value to set upon these remarkable results, in the absence of 

 information as to what exactly was the plant used by Bell Salter 

 under the name tetragonum. If it were adnatum Griseb., his 

 results conflict with Haussknecht's and mine as regards the 

 complete sterility of the hybrid. If it were obscurum Schreb., 



