TUE Sl'KRM.VTOGENESIS OF DOilESTIC MAJiM.VI.S. 243 



it has beeu deseiilHil l>\ Wüdsbdalek ('14) in the Lorse, BACHiiVBEn ('16) iu 

 tlio mbbit nufl Allen ('18) in tho nit. Li liistn-ts this ]y>i]y W!i3 accumtely 

 describecl by Montgomery ('11) und "SVilsok ('13). 



l'iit its orif^iu nud functiun jire still n, question. It is alrciidy knu^'u ihi-A. 

 iu thi; siieraui,tid of the monse (C. Niessip;') and oi" tho rat (Meves) this Ixxly 

 is very conspicuous, while iu the guinea-pig (Meves) it disappears at the 

 lieginning of the development of the spermatozoa. Meves ('99) in the guinea- 

 pig showed tlrnt the chi-omatoid body is sfay'npxi as the nucleoli. He aiys : " Ich 

 selbst LiIks von Färbungen, welche geeignet sind, über die Natur des cliroma- 

 toiden Nolxjnkürixjrs Aufechluss zu geben, mu' (nach Sublimatfixirung) die Ehi'Uch- 

 Biondischo Di-cifeichfärbiujg angewandt, lx;i der er sich ebenso vde die Nukleoleu 

 intensiv rot färbt ; ich kann also jedcnfidls Moore nicht beistimmen, diiss es 

 sich um eliminertcs Chromatüi hsmdelt." Duesberg ('11) also in tho guinea- 

 pig found the same phenomenon by using Besd.v's method. Allen's ('18) 

 observation in the rat is different from that of many other iuvestigiitors as well 

 as from mine. His description on its behavior and on its £ite is as follows :" Its 

 Erst pisitive apjiearance is in the late propliase stage of the fii-st Sjiermatocj'tes 

 after the disappearance of the nuclear membrane. At fii-st it appears to lie 

 near the chromosomes, but at a later stage it is always found well out in the 

 cytoplasm. In some meta,plmse cells it is doubled. It is lost dm-ing the 

 amipliase of the first siiermatocytes and during the iuterkinasis stage, but reap- 

 l^eais in tho second spermatocytes. Nothing of ecjuivalent form is foimd iu the 

 spermatids. In these cells, however, there is a mass lying near the nucleolus 

 which stains Kke chromatin. It develops intensity of staining reactions as the 

 siTcrmatids advance iu differentiation. Li its fuller development it is as in figiu'e 

 1, si)ermatid 3, where it ap^x^ars as a globular body, but much Lirger and 

 staining more deejily than the chi-omatoid bodj'." 



In my materials, as stated alx)ve, the chromatoid lx)dy is coaspicuous from 

 the prophase of tha fiivst division up to the transfonnation of tlie spermatids, 

 and as Meves ('99) and Ddesberg ('11) indicated, it stains deeply with acid- 

 fuclisin \nth Auerbach 's method. As to the fate of this body there is s, 

 diftereuce between the mouse and the rabbit. 



In the moase it remains near the centi-osomes within the " St-hwanz- 

 manschette " during the development of the spermatozoa, while in the latter it 



