NOTES ON THE NOMENCLATURE OF FUNGI 79 



From this it will be seen that the earHest specific name after 

 Fries's Systema is " quadrata," and on the grounds of priority 

 alone a new combination would have to be made. This kind of 

 thing occurs in other genera of the Phycomycetes. Eeferring to 

 Art. 49 bis of the International Eules ='^ it will be seen that 

 Phycomycetes are not mentioned, and from the wording (which is 

 open to criticism on other grounds) it would appear that they were 

 not even considered. In the case of Syzygites and Sporodinia, the 

 spore forms are so distinct in appearance and in origin that they 

 received different generic and specific names ; there can be no 

 hesitation in considering the zygote to correspond to what it has 

 been agreed to call the " perfect form " and to have therefore the 

 prior right to a name. Many Oomycetes were given names in 

 their conidial stage but in the great majority of cases the oosporic 

 stage was not discovered until the doctrine of pleomorphy was 

 becoming firmly estabhshed, and hence did not receive a new name 

 although the two kinds of spore are very different. If the oospore 

 had received a different generic (or even specific) name, that 

 name would clearly have to stand. In the special case under 

 discussion the oospore stage was first described by de Bary, 

 and as it was recognised to be the same fungus as that of 

 which the conidial state had received the name Gystoims,^ the 

 same name was used. In other words the name of the conidial 

 stage is Cystopus Lev. and the name of the oosporic (or perfect) 

 stage is Cystopus de Bary. Therefore the specific names must 

 begin with de Bary. 



This is the logical way of looking at the question, and it has 

 the additional merit of retaining the majority of the names in 

 common use in the Phycomycetes owing to the fact that a know- 

 ledge of the sexual stage was gained principally by the efforts of 

 Tulasne and de Bary, and the names used by these distinguished 

 investigators were naturally generally adopted. Thus de Bary's 

 names, Cystopus candidus, G. Lepigoni, C. Portulacae and C. Bliti 

 have been generally used by British mycologists. The remaining 

 species usually known as C. Tragopogonis should be called 

 C. cuhicus. 



Cystopus cubicus Lev. emend de Bary in Kabenh. Fung. 

 Europ. No. 480 (1862) ; Ann. Sci. Nat. 4th ser. xx, 132 

 (1863). 



Uredo Candida Pers. f3 Tragopogi Pers. Syn., 223 (1801). 



U. obtusata Link in Mag. natarf. Freunde, Berhn, iii, 6 

 (1809). 



U. cubica Strauss, Wetterau. Ges.f. Naturh. ii, 86 (1810). 



U. Tragopogi DC, Fl.fr. ii, 237 (1815). 



Albugo Tragopogi S. F. Gray, Nat. An: Brit. PI. ii, 540 

 (1821). 



Uredo Candida b. compositarum, Eabenh. Kryptfl., i, 13 

 (1844). 



Uredo (Cystopus) cubica Lev. in Ann. Sci. Nat., 3rd ser., 



viii, 371 (1847). 



* Cf. Journ. Bot. 1915, 302. 



