42 Journal of Agricultural Research voi. xv, no. i 



results, it is entirely likely that their presence would be noted because 

 the pattern of each is so striking. It does remain to be shown rather 

 that the piebald cattle, like the Shorthorn, have a difference in behavior 

 of the separate spots which compose this piebald. A beginning at this 

 kind of analysis has been made by Kiesel, according to a review by Lang 

 {i8). In these experiments a solid-color Limburger race was crossed to 

 a piebald race, F^ intermediate piebald. The back cross gave 22 solid 

 color and 29 piebald. The back cross to the piebald gave 84 piebald out 

 of 90. Unfortunately, no record of the exact spotting has been given; 

 consequently, we are left in the dark concerning any difference in behavior 

 throughout the coat. It would seem, however, that Kiesel's results 

 would fall in line with the results obtained here, where each individual 

 area is treated separately. 



Analysis by the method of individual white spots has shown a marked 

 difference in the inheritance of such spots throughout the animal's coat. 

 This is perhaps as far as the writer should go, and is the only conclusion 

 it is intended to emphasize; but realizing that there are exceptions not 

 yet accounted for, it may be said that white spotting in the inguinal 

 region is, broadly speaking, dominant. The spots of the rest of the pie- 

 bald pattern are, individually considered, recessive. These enumerated 

 individually according to the region in which they occur are (i) white on 

 the face (star, star snip, or blaze) ; (2) white on the throat; (3) white as 

 a band across the shoulders; (4) a white area on the rump at the base of 

 the tail set ; (5) white on the flanks as irregular spots ; (6) white on the 

 tail above the switch ; and (7) the white stockings on the four feet. 



The bearing of this difference in inherited behavior on the general 

 problem is at once evident. If the red-coated Shorthorns should carry 

 one of these recessive white spots, we should expect a small proportion 

 of cattle produced from the random mating of such an animal in the 

 Shorthorn population which would be white-spotted. In point of fact, 

 this is what has actually been obtained. The reverse is also true that if 

 these dominant white are mated together, we should expect that a het- 

 erozygous mating would now and then take place, giving a red. The 

 evidence brought forward offers a straightforward, clear explanation of 

 the anomolous behavior of the Shorthorn coat. 



Behavior of the Muzzle Color in the F, Progeny 



The categories used to describe the color of the muzzle pigment are, 

 in general, the same as those used for the tongue color. The tabulation 

 and classification of these terms has already been given by Pearl (33). 

 For this classification the reader is referred to his paper. In his study of 

 tongue color it was found that the essential thing was not so much the 

 color of the pigment but rather its presence. This study will, therefore, 

 be limited to the presence or absence of pigment. Table XXHI gives 

 the tabulated data for this treatment. 



