538 



Journal of Agricultural Research voi. xix, no. n 



and lax heads. Thirty-seven produced progeny consisting of types 

 with inclosed emmerlike kernels, intermediates for thrashing, and 

 common plants. Of these 37 F3 lines, 17 produced progeny of only two 

 sorts, emmerlike for thrashing and compact keelless (PI. 100). 



It is, of course, impossible to determine the actual factors involved in 

 this cross, because some gametes or zygotes were eliminated by sterility. 



The cross in which emmer (C I 1524) was the female parent gave results 

 similar to those obtained when emmer was used as the male parent. 

 This is shown in Table IX. One family, 169-5, which was grown from 

 an intermediate F, plant, produced emmers, intermediates, and common- 

 headed sorts and was quite resistant (PI. 100). 



Table IX. — Classification of crosses between emmer {Minnesota Il6^ and C I 1524) "with 

 Marquis on the basis of rust class in the F^ generation as determined by the Fo '^'^'^ ^a 

 generations , and the m,ain character differences separating emmer from common wheats as 

 determined by the Fj and F3 generations 



MARQUIS X EMMER I165 



EMMER (C I 1524) X MARQUIS 



It is interesting that out of a total of 73 plants from these two crosses, 

 8 bred true to the emmer habit for thrashing and for head shape. Of 

 these 8, 7 plants were put in rust class 1 while i was practically resistant, 

 the progeny in the F3 generation being placed in rust class 2. Only one 

 lax, common plant was obtained in the Fj generation, and this was sus- 

 ceptible. Several plants were obtained in the F3 generation which were 

 not only rust-resistant but also resembled common wheat. 



