PE3I0ME 711 



II opraHOB aapo^biuia. 3to Ka>KeTCH tcm 6oJiee CTpaHHbiM, hto B. Py HSBecTCH kbk 

 i),iiiH H3 Hpbix aainHTHUKOB ii/ien o cymecTBoeaHHH CBaaii Me>K;iy cjjopMOH ii (jjyHKUHCH. 



OaHaKo Ha ocHosaHHH flnajieKTHHecKo-MaTcpnajincTnqecKoro yiewHH 06 OHToreHern- 

 qecKOM pasBiiTiiii HejTb3fl AonycTHTb cytuecxoBaHUH npeAcjjyHKUHoHajibHbix 4^33, b reneHHe 

 KoTopbix pasBHTiie 6bijio 6bi oT^ejieHO OT c|)yHKmiii. 



lima HCTOMHiiK npimiiH iiH;inBHAya.fibHoro pa3BiiTiia bo B3anM0AeftcTBiiHX, pasBii- 

 BaK)LUnxcH B OHToreHeximecKoM KOMnjieKce, 11 cqHxaH 3c{)c})eKT 3thx B3aHM0AeHCTBHH KaK 

 npoHBjieHiie peaKTHBHoCTH JKneoro semecTBa (axa peaKXHBHocxb H3MeHnexcH KojiimecxseH- 

 110 n KaMecxBCHHO na pa3Hbix cxynennx oHxorenesa), HejibSH oxAejiiixb cjjyHKnmi SMopno- 

 Ha.ibHbix saMaxKOB ox BsaiiMOAeiicxBHii, Koxopbie 3xh (|)yHKmin no;i;iep>KHBaKDX Me>KAyHHMii. 



KaK y>Ke 6bi.rio cKasaHo, cymecxByex nocxoHHHoe pasBuxne 3xiix B3anM0AeHCXBHH 

 II, cjieAOBaxejibHO, cjjyHKUiiH 3M6pnoHajibHbix opraHOB na pa3Hbix cxyneHHX pasBHXHa, ko- 

 xopoe Be;iex b pasHbix /ijih Ka>K;ioro oprana (h ana Ka>Kaoro sHAa) nepHoaax K xaK na- 

 3biBaeM0My «BspocjlOMy cjjyHKmionajibHoMy nopory», xo ecxb k noHBJieHHio cjjyHKUiift, cne- 

 micjjiiMecKiix R.nH BnoJTHe pasBiixoro opraHa. 



Cjie/iOBaxejibHO, «B3pocjTbiH (jjyHKUHOHajibHbiH nopor» 3X0 ne norpaHiiHHaa MexKa, 

 ox^e-UHiomafl oahh c})yHKUHOHa.xbHbiH nepHo/l ox apyroro (J)yHKUHOHajibHoro nepno/ia. a 

 KpHXHWecKaH xo^Ka (^yHKUHOHajibHoro Ka»jecTBeHHoro H3MeHeHHH. 3xh KpHXH^ecKiie mo- 

 MCHXbi, CBHsaHHbie c ii3MeHeHHaMii xiina o6MeHa Bemecxe, Haxo/iHXca bo MHorHx cjiy^aax, 

 KaK HanpuNiep y KvpiiHoro 3apo/ibiuia, b cooxHouieHHH c xaKiiMH asjieHHaMH, KaK pa3Biixne 

 /KeJixoMHoro KpoBoo6paui,eHiia, nepexoA k reiMaxosy nocpeiicxBOM ajijiaHxoiicHoro KpoBoo6pa- 

 meniia, o6pa30BaHiie noAsapoabiujesoii >khakocxii nyxeM nepeMemeHna Bo^bi 113 6e.iKa b 

 «ejixoK H ap. 



3M6pH0HajibHbiH oHxorenes npeacxaBJiaex xaKUM o6pa3oM pa^ nepexo^Hbix nepwo- 

 AOB, B xeieHiie Koxopbix npoiicxo/iax ii3MeHeHiia ycJioBHH pasBnxiia. 



3th nepexoflHbie nepnoiibi Hsynaioxca He xojibKO paAii hx snaMeHna b iiccjie/ioBaHHii 



paSBHXllH (j)yHKU.HH 3M6piIOHa.îIbHblX OpraHOB, HO H AJIH XOTO HX06bl 3XHM HyXCM HafixH 



(b CBexe y^eHHa B. MHiypHHa) cnocoSu nanpaBHXb xe^eHne oHxoreHe3a no hobbim nyxaivi, 

 ii3MeHaa ycjioBHa pasBHxna b xe^eHHe sxhx nepHOfloB. 



MimypHHCKoe yneHiie AonycKaex B03MO>KHocxb paciiiaxbiBaHiia nacjieAcxBeHHOcxii 

 npii noMomii xaKiix Biweiuaxe.ribcxB. 



Hejibsa flonycxiixb cymecxBOBanHe cjjyHKUHonajibHbix mjih npefl(})yHKmiOHajibHbix 

 3M6pHOHajibHbix opraHOB. JXo ;iocxH>KeHHa B3pocjToro Mopcjjo-cJDyHKUHOHajibHoro nopora 

 3M6pnoHajibHbie opraHbi Moryx HMexb BsaHiviooxHoiueHiia h c[)yHKiuiH, ox.inHaioiuiieca ko.fiii- 

 HecxBeHHO H KanecxBeHHO ox KoneHHbix. 



3M6piioHa.iibHbiii opraH aBJiaexca Bcema Heo6xoAHMbiM pe3y.TbxaxoM B3aHM0Aeii- 

 cxBHH Bcero 3M6piioHajibHoro KOMnjieKca 11, b cbok) onepe/ib, cjjaKxopoM b 3xom kom- 

 njieKce, npoHBjiaromHM xaKHM o6pa30M cboio (j)yHKUHio. CjieAOBaxeJibHO, Ka>KAbiH 3M6pH0- 

 na.xbHbiH opraH npoxoanx HecKo.ibKO c})yHKHHOHa.TbHbix (J3a3 JXO ooaBjreHHa cnemicJjHHecKoft 

 ijn'HKUHii, xo ecxb cjjyHKUHH, no;io6HOH cyinecxByK)m,eH bo BnojiHe pa3BHX0M opraHn3Me. 



OyHKiuiii, Heo6xo/iHMbie jijih «iisHeAeaxejibHocxH h pa3BHXHa 3apo/ibiuja, Moryx 

 HMexb flBa poaa Mop(})ojiorimecKiix cy6cxpaxoB : 



1. OpraHbi, HMeiomne yHiixapnoe pa3Biixiie, Koxopbie ocxaioxca xaKOBbiMii ox naMajia 

 JXO KOHua OHxoreHe3a, nocxoaHHO npHcnocaâjiiiBaacb k Hy>KaaM SMopiioHa.ibHoro kom- 

 njieKca, KaK nanpiiMep : cep/iue, ne^eHb. 



2. KoMHJieKCbi opraHOB, npeAHasHaqeHHbie rjmi BbinojiHCHiia cxoAHbix i|)yHKUHH, ho 

 npe^cxaBJieHHbie xonorpacjjimecKH oxAejibHbiMH KoivinoHeHxaMH, Koxopue jxoxojxm jio hx 

 cnemicl)HiecKOH cj)yHKUHOHaj:bHOH (jjasbi b pa3Hbie cxa^HH oHxorenexHHecKoro npo- 

 uecca, npHHaiiJie>Kamiie, o^HaKO, oiiHOMy h xoMy >Ke cjjyHKUHonajibHOMy KOMOjieKcy. 



3xH 3M6piioHa.ibHbie opraHbi Moryx obixb Ha3BaHbi roMeocjjyHKUiiOHajibHbiMH b xom 

 CMbic.ae, Mxo ohii npe/inasHaieHbi /iJia iicno.THCHHa oflHofl n xoii /Ke (}5yHKmiH. 



KaK npHMep roMeocjjyHKUiioHajibHofl napbi mo>kho yKa3axb xopHoajiJiaHxoHC h jierKiie. 



Ha^HHaa c 8-ro ^na iiHKyoauHH r;iaBHbiM jţbixaxejibHbiM opranoM nxHHbero 3apo- 

 Abiuja aBJiaexca xopiioa.rr.iraHxoHC. 0;iHaK0 oh cocymecxByex c ApyrHM abixaxejibHbiM 

 opraHOM, a iiMeHHo jictkhm, Koxopbiii ;iH(|){j)epeHHHpyexcH MCfljieHHO b xeieHne 3M6pHo- 

 HajibHoro oHxorenesa 11 aoxo^HX jio cBoefi «Bspocjioft» cneu,H(f)HMecK0H ctiyHKuiiii HeMHorn 



