742 HCCJIE^IOBAHHH nO 3KCnEPHMEHTA:ibH0n 3MBPH0JI0rHII 



paHbiue BbijrynjieHHH, Kor;ia aapoabim HaMHHaer Abiuiarb BoaayxoM h3 B03;iymH0H KaMepu. 

 B 3T0T MOMeHT HamiHaeTCH o6paTHoe pa3BHTne xopHoajiJiaHTOHca. 



B npuBefleHHOM npHMepe o6a naprHepa roMeo(j)yHKUnoHajibHOH napbi 3to paajiimHbie 

 opraHbi, TonorpacJ)HMecKii OTiiajieHHbie, oanH BHyTpiisapo/ibiujeBbiH, a ^pyroH BHesapo- 



flblUieBblH. 



B cjiynae nepBHMHofl h ETopuiHoii noMcK (mcso- ii MeTaHe(J)poc) Ha6;iK3;iaeTcn 

 nocjieAOBaTejibHOCTb (bo BpeineHii h b npocrpancTBe) Tonorpa(J)iiMecKH pasaejibHbix hoch- 

 TCJieH BbiAe;iiiTejibHbix cj)yHKUHH. O/iHaKO s^eob nocjie/toBaiejibHocTb npoHcxoflUT paHbiue, 

 B TeneHne SMgpHOHajibHoro OHxoreHesa, nponcxo;inmero ;io BbijiynjreHiiH, a o6a napTHepa 

 napbi npnHaflJie>KaT 3M6pHOHajibHOMy tejiy. 



HesaBiiCHMO or Toro, 6yaeT ;in Bcn He(})poreHHaH Macca (npo-, Meao- ii MCTaHC- 

 (j)poc) paccMarpiiBarbCH KaK eanHbifl «ro.TOHecjjpoc», nocjieflOBarejibHo ;iH4)c})epeHUHpyio- 

 miiflcH II a^anTHpyiomHHcfl b KpaHHo-Kay;iajibHOM HanpaBJieHHH, iinn Bce rpH noKo.aeHHfl 

 noHCK 6y/iyT paccMaxpHBaTbCH KaK caMOCTOHrejibHbie oSpasoBaHHH, b o6ohx cjiynaHX 

 6y;iyT Ha6j]K);iaTbC5i — b oTjiHiHe ot neweHii, cepaua ii t. a. — pasflejieHHe bo BpeMenii 

 H B npocrpaHCTBe opraHOB, HcnojiHflK)mHX noflcSHbie cJ)yHKUnn h wepe/iyioimixca b npo- 

 uecce oHToreHeximecKoro pasBUTHH. 



BawHO, oflHaKO, xo, mto Meao- h MeTaHe(})poc He saiviemaioTCH crponiM o6pa30M h 

 cocymecTByror o;iho BpeMH, oSpaayn o;iHOBpeMeHHO napy, cocTonmyK) h3 BbiaejiureJibHoro 

 oprana, KOTopbifl naxoAnxcn b pa3rape iiejiTejibHOCTH h npHcnoco6jieH k aKiyajibHbiM 

 TpeSoBaHiiHM 3M6pnoHajibHoro KOMHJieKca (Me30He(J)poc), H ;ipyroro opraHa (MeTaHe(j)poc), 

 KOTopbiH cymecTByer Tonorpac})HHecKii pasaejibHO ot nepBoro, ho eiue He ^ocxnr CBoeii 

 cneuH(J3HHecK0H aH(})(})epeHmiaunH. SareM 3Ta napa npoxoflHT h (})a3y cneuH(j)H»iecKofl 



CJ)yHKU,HH, 0/(H0BpeMeHH0H jinn 06OHX napTHCpOB. 



Oâ-beKTOM H3y«îeHHH 6bi;ia n36paHa roMeo(})yHKUHOHajibHafl napa, onpe;ieJieHHaH b 

 BbiiueyKaaaHHOM cMbicjie, a HMeHHO napa Me30- h Meranecfjpoc, xaK KaK, no-BHflHMOMy, 

 MOMnjieKC TaKoro xnna SMâpHonajTbHbix opranoB oâjia^iaex ocoSbiMH npeHMyiaecTBaMH RJifi 

 3KcnepHMeHTajibHoro HayHeHHn no OTHOuieHHKD k «efliiHbiM» opranaM tuna cep^ua, 

 neieHH h t. a- 



3KcnepHMeHTnpoBaHHe c xaKofi napoft opranoB Morjio 6bi npHBecxH k Bbipa6oxKe 

 MexoanKii flJiH aHa;iH3a OHXoreHHiecKux npoueccoB, Koxopue cnoco6cxByiox aH(J5(})epeH- 

 miauHH 3M6piioHajTbHoro oprana b npHcyxcxBim ero roMeo^jyHKUHOHajibHoro napxnepa — 

 eme Bno;iHe Aocxaxo^Horo ajifi xpe6oBaHHH 3M6pHOHajibHoro KownjieKca npeauJecxBeHHHKa, 



To, 1x0 B nayMaeMOH nape pa3JHiMHbie Mop(})o-cJ}yHKmiOHa;ibHbie oHxorenexHiecKHe 

 (Jiasbi BcxpeiaroxcH o^HOBpeMeHHO b pasHbix, Tonorpac})ntiecKn pa3;ieJieHHbix opranax. 

 npeacxaBJiaex Sojiee Bbiro/iHbie ycjiOBiiH jiJifi 3KcnepiiMeHXiipoBaHiiH. 



B 3X0H rjiaBe ii3yiaK)TCfl MOpcJ)o-(})yHKU,HOHajibHbie ocoSenHOcxH nepBiiMHoft ii bxo- 

 pimHOH noweK b xeneHne 3M6pnGHajibHoro onxoreHeaa npH noMoiuii HecKOJTbKiix iviexoaoB, 

 cnocoSHbix BbiHBHXb (})yHKmiOHa;ibHbie CBOHcxBa 3XHX flByx noKOJieHHÎi noweK. 



Ope/inpHHHXbie HccjieiioBaHiiH nosBOjiHioT yxBepwaaxb, hxo cyuiecxsyrox neonpo- 

 Bep>KHMbie ;ioKa3aTejibcxBa cneuH(})HHecKOH (J)yHKU,Hii b MexaHe(})poce 3apoAbiuja 6-flHeBHOH 

 HHKySauHH. CxonjieHiie xpHnaHOBoro cuHero b KJiexKax nsBiixbix KanajibueB Me30He4)poca 

 b 3xoii cxa^iiH OTMeiaex cymecxBoeaHHe cneun(j)HiecKH noieiHOH (})yHKUHH ; iiaBecxHo, 

 MTO CKonjieHHe xpHnana xapanxepno jijm H3BHXbix KaHajibueB no3BOHOMHbix. ripucyxcxBiie 

 c(})epojiHxoB MOweBOH KiicJTOxbi iijiJiiocTpHpyeT HecoMHeHHoe BbiiiejieHHe SToro BemecTBa, 

 HawHHaH c 6-ro jxhh iiHKyGauHH. 



HajiHHiie me,JioqHoii (})0C(j)aTa3bi b aniiKajibHOH o6ojioMKe kjictok hsbhtux KanajibueB 

 H rjiHKoreHa b o6jiacxn anuKajibHoro h 6a3ajibHoro nojirocoB xex «e KJiexoK ;iOKa3biBaex, 

 qxo c 5-ro mu HHKySauHH kjicxkh HSBHxbix KaHajibueB BbinojiHHiox (})yHKUnio BcacbiBaHHfl 

 rjiroKosbi H AenoHHpoBaHHH ee no;i bhaom r.miKoreHa. 



3xox nepHo;i (5 — 6 ;iHeH HHKy6auHH) coBnaAaex c noHBJieHHCM cnoHxaHHofi ^'•'lyo- 

 PeCUeHUHH B Me30He(J)pOCe, KOXOpyro MOJKHO BUHBHXb h iisyMHXb npH noMomii CJJHJlbXpU- 



PoBaHHbix yjibxpa(J)HOJiexoBbix jiyMefl (By;i). 



TaKHM o6pa30M na 5— 6-h iienb HHKy6auHH Me30He(j)poc oâjia^aex neocnopHMO xapaK- 



XepHblMH nOMeMHbIMH {jjyHKUHHMH. B 3X0M nepHO;ie Me30He(})pOC B COCXOHHHH lICnOJIHHXb 



cBoii cnemi(})niecKiie (|)yHKmiii. Sxiim (jjynKUHîiM cooxBCTCTByeT ii Kapxiina xonapnoivia, 



