134 PRIMATE ALVEOLINGUAL SALIVARY AREA 



nerve does not bear the same relation to the lateral alveoUngual 

 field. The details in the carnivore type are further considered by 

 Schulte (Part VI). 



Consequently, in the primates, the individual glandular anlages of 

 the entire available alveolingual field, considered as originating at 

 ecjuidistant intervals from each other, will encounter secondary modi- 

 fying influences in the cephalic, intermediate, and caudal divisions of 

 the entire area. 



Cephalad of the intersection with the lingual nerve the topographical 

 and mechanical relations of the alveolingual gutter will permit of a 

 relatively free unfolding of the primitive anlages. At the level of the 

 lingual intersection the entire available alveoUngual area will experi- 

 ence a transverse constriction of the potential glandular field, and tend 

 to tilt its individual lateral components dorsomesad toward the faucial 

 and palatine areas, while its medial components will already tend to 

 escape from a strictly intermandibular position, above the mylohyoid, 

 in a caudolateral direction, beyond the confines of the caudal margin 

 of the mylohyoid into the far more available submaxillary interval 

 between this muscle and the digastric. This reconstruction of a pos- 

 sible phylogenetic path of intermandibular glandular development is, 

 perhaps, of value in explaining the intersection of the drainage lines 

 presented by the adult mammalian types of separate and distinct ducts, 

 with indifferent separate glandular development along the lateral 

 border of the primitive available field. 



It seems reasonable to assume, without direct reference to the ontog- 

 eny of the individual glands presently to be considered in detail, that 

 the escape from the narrowly limited confines enables the primitive 

 glandular anlage to develop more fully into the secretory apparatus 

 of the adult, while the mechanically more limited and restricted area 

 of the primitive generalized field will, under these" conditions, suffice 

 for the estabhshment of a duct, a structure necessarily less exacting 

 in its demand on the available space, compared with the bulk of the 

 secreting gland, and capable of utilizing in addition any remaining 

 portions of the adjacent primitive field for the development of acces- 

 sory and secondary secreting glandular collections. 



The impression which a comparative study of the adult mammalian 

 intermandibular salivary complex gives is that the relation between 



