Volume I NOVEMBER, 1911 No. 4 



Nr;\V Yoj^ 



ON GAMETIC SERIES INVOLVING REDUPLICATION *"" ' ^ "^'^ac 

 OF CERTAIN TERMS\ "''"'"'' 



By W. BATESON, M.A., F.R.S. 

 AND R. C. PUNNETT, M.A. 



In a paper recently published- we gave a brief account of some 

 peculiar phenomena relating to the coupling and repulsion of factors in 

 the gainetogeiiesis of the sweet pea and of several other plants. The 

 view there stated was that if A and B represent two factors between 

 which coupling or repulsion can exist then the nature of the F., 

 generation depends upon whether A and B were carried into the 

 Fj heterozygote by the same gamete or by different gametes. If the 

 heterozygote AaBb is formed by the gametes AB and ab partial 

 coupling between A and B occurs in F.^ according to a definite system, 

 and it must be supposed that the gametes formed by the heterozygote 

 belong to one or other of the series 



•3AB : Ab : aB : 'Sab, 

 7AB -.Ab-.aB: Tab, 

 loAB -.Ab-.aB: \5ab, &c. 



If on the other hand the heterozygote, AaBb, is formed by the gametes 

 Ah and aB repulsion occurs between A and B, so that only the two 

 classes of gametes Ab and aB are formed. In the account to which 

 we have alluded we supposed that such repulsion was complete, and 

 that the two classes of gamete AB and ab were not formed. Our work 

 on sweet peas during the present summer has led us to modify our 

 conception of the nature of the gametes produced in cases where 

 repulsion occurs, and this modification will perhaps be made clearer if 

 we begin by giving an account of the experiments upon which it is 

 based. 



' This papeV is also appearing in the 49th volume of the Briinn Verhandlungen which 

 is to be published as a Mendel Festschrift. 

 2 Proc. Ray. Soc. B, Vol. 84, 1911, p. 1. 

 Journ. of Gen, i 20 



