A. R Blakeslee 19 



little more than the mid-ribs. If the Globe mutant, which has broad 

 leaves, becomes Q., the leaves become narrower than typical Globes, 

 but are still broader than normal narrow-leaved types. An increased 

 indentation, however, is characteristic of all Q. leaves, whatever may 

 be the genetic type to which the plant belongs. 



The stature of Q. plants is somewhat less than that of normals. 

 The branches are more slender and the root system is less well 

 developed. 



Typical Q. flowers are recognizable at an early stage. A series with 

 buds is shown on PI. Ill, fig. 3 in comparison with normal flowers below. 

 The corolla of Q. flowers is split between the lobes as far down as the 

 insertion of the filaments as shown in the opened flower on the extreme 

 right from which the calyx has been removed. The corolla segments 

 are incurved and twisted around the base of the flower. Frequently in 

 the expansion of the bud they are unable to free themselves from the 

 calyx. The third and fourth Q. buds from the left are typical of such 

 " blind " flowers which may fall from the plant without opening. The 

 infolding of the lobes is characteristic of early stages, and hence, when 

 a normal bud would have its corolla nearly reaching the tip of the bud, 

 the incurved lobes leave an empty space inside the apex of the calyx 

 at the stage shown by the second bud from the left. The flowers of 

 purple Quercinas are darker coloured than normals and the stem colour 

 in such forms is also increased in intensity. 



The stamens in the bud (Fig. 4) are found to be shrivelled, and 

 in typical Q.'s produce nothing more than a few small grains which 

 when tested have been found to be functionless. 



The stigma in normal plants is made up of two thick lobes, facing 

 outward (Fig. 4). In Q. flowers the stigmatic surface is chiefly on the 

 inside of the lobes and runs part way down two sides of the style. 



The most conspicuous peculiarity of Q. plants, however, is the 

 suppression of spines on the capsule. This may be complete, giving 

 capsules as smooth as those of the inermis variety. The form of the 

 capsule, however, is that characteristic of the variety or mutant aflected, 

 the globe mutant plants, if also Q., having flattened globose fruits. 

 Seeds from Q. capsules are distinctly smaller than those from normals 

 and their percentage of germination is less. The cotyledons are fre- 

 quently unable to extract themselves ft-om the seed coats without 

 assistance and plants may die for this reason. Older seedlings also 

 appear to be less vigorous than normals. 



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