A. F. Blakeslee 29 



Naudin (I.e., p. 49) reports in an F^ between an inermis variety 

 (7). laevis) of D. Stramonium, and the armed type, that while most of the 

 plants had spiny fruits, others had fruits with reduced spines. Many of 

 the capsules on three out of the 40 plants in this generation were 

 very spiny on part of the surface while totally smooth on the rest. 

 Naudin believed that they united thus by distinct and separate com- 

 partments the distinctive traits of the parental types {D. laevis and 

 D. Stramonium). He calls this " liyhridite disjoints " and cites in this 

 connection the condition in the graft chimaera Cytisus Adami. He 

 attempted to secure offspring from the smooth part of such a mosaic 

 capsule, but due to the poor maturity of the seed only four seedlings 

 were obtained. Of these, one was inerm,is, and may have been an 

 extracted recessive or a strong Q, In a series of F2 plants from the same 

 original cross, Naudin found six individuals out of 38 which again 

 showed more or less well marked his " hyhridite disjoints " and which 

 were also presumably Q.'s. 



Godron (I.e., p. 14) reports finding capsules partly spiny and partly 

 smooth. He objects to Naudin's interpretation that the separation of the 

 fruit into smooth and spiny portions is due to their origin from smooth 

 and spiny parents, since he says that he has found this condition when 

 both parents had spines. 



Bateson and Saunders (I.e., p. 23) after discussing the intermediate 

 colour of the flowers of the Fi between white and purple flowered forms 

 say: 



" The occurrence of intermediate forms was also occasionally notice- 

 able in the fruits. Among the large number of capsules examined, there 

 were some of the mosaic type, in which part of the capsule was prickly 

 and the remainder smooth, while others, suggesting a blend, were more 

 or less prickly all over, but the prickles were much reduced in size, and 

 often formed mere tubercles. These mosaics occurred as rarities both on 

 prickly individuals and on smooth ones still more rarely." 



Further evidence pointing to the Q. nature of these abnormal cap- 

 sules is given in their following statement : 



" Such intermediate fruits were most often found towards the end of 

 the flowering season." 



In a footnote (I.e., p. 24) they call attention to a plant with a single 

 undersized smooth fruit in the Fi from the cross, purple inermis x white 

 armed, where onl}'^ armed plants heterozygous for inermis should be 

 expected. If their suggestion of possible experimental error for the 

 appearance of this plant be not the correct explanation it may have 



