MILIUSA AND SACOOPETALUM. 163 



the appendages of the corolla appear as tufts of hairs inserted at the 

 base of the torus, remaining after the petals have been taken away. 

 It should be here noted that the whorl which the author described 

 as a calyx may have been in fact an outer series of petals, conform- 

 ing, as in certain allied species, to the true sepals, which in this case, 

 however, must have been highly caducous. 



In 1834 Wight and Arnofct (Prodromus p. 10) under " Miliusa 

 indica (Lesch.) " reproduced A. Le Candolle's leading characters, but 

 assigned them to " Wall. L.n. 5433, " quoting as a synonym 

 Uvaria ciliata, Heyne MSS." There is a specimen at Kew ticketed 

 Uvaria ciliata, Heyne Tenmalej near Courtallam, July 1815," and on 

 the same sheet is a second ticket from which it seems probable that 

 the citation of " Wall. L.n. 5433 " may refer to this specimen or to 

 a duplicate. 



In April 1835, Wight got a plant at Courtallam which he ulti- 

 mately referred to Miliusa indica, and in the following July and August 

 he collected further specimens which together with the April gather- 

 ing have since (1866-67) been distributed from Kew as No. 34 of the 

 Indian Peninsular Collections. These later gatherings (in part) 

 resemble " Uvaria ciliata, Heyne " so far as can be judged, but 

 Heyne's specimen is not in fruit : No. 34 of Wight has drupes and 

 matches an example from his own Herbarium, without date or original 

 locality, with which is now attached a description in his own band- 

 writing. This description, however, seems to have belonged to the 

 specimen distributed under No. 33 as " Miliusa montana, Gardner " 

 (which in PL Brit. Ind. i. 86 is reduced to M. indica) and to have been 

 pinned to the sheet where it is now found by accident. A pencil note 

 on the sheet from Wight's own Herbarium (in Sir J. D. Hooker's 

 handwriting apparently) points out that the plant there represented 

 differs from No. 33 by the ripe carpels being sessile and pubescent- 

 tomentose. This form (No. 34 in part) seems to have received from 

 Wight the manuscript name of " M. affinis ". Nothing on these sheets 

 can be referred to Alph. De Candolle's M. indica unless we assume — 



(1) that the sepals, which in Nos. 33 and 34 Herb. Pen. Ind. are 



manifest, may sometimes fall off at a very early stage, or 



(2) that the sexes are diclinous and that the structure of the an- 



droecium differs in the male and female flowers very widely, or 



(3) that the arrangements of the floral whorls is remarkably 



unstable. 

 How far any parallel to the degree of instability that must be 

 supposed in this case has been observed in other Anonaceaa is a ques- 

 tion which may be deferred for the present. In the " Illustrations " 

 i, p. 68 (1840), Wight met this difficulty by suggesting that Da, 



