562 



With regard to internal characters of less importance I may men- 

 tion there is no gizzard in the anterior segments, but five gizzard 

 like structures one to each segment, beginning with the XVII.; there 

 are a large pair of calciferous glands in segment XIII. as in 

 Euclrilus and tAvo unpaired »Chylustaschen« arising from the ven- 

 tral surface of the oesophagus in the XI, and XII. segments. As in 

 Shihlmannia these organs are richly supplied with bloodvessels ; their 

 epithelium is folded, and the folds have become united so as to give 

 the appearance of a series of parallel tubes: the distal extremity of 

 each pouch is remarkable for the fact that the ductules have become 

 in tra -cellular and form an anastomosing plexus which presents a 

 striking ressemblance to a nephridial network. 



The structural characters of more particular interest concern the 

 reproductive organs. 



The testes appear to be normal; the sperm sacs are paired and 

 lie in segments XI., XII. attached to the front wall of each of these 

 segments. As in Teleudrilus (see Rosa »Lombrichi dello Scioa« Ann. 

 Mus. Civ. Genova, Ser. 2. Vol. VI) the funnels of the four vasa 

 deferentia lie in the interior of the spermsacs; each funnel is con- 

 nected by a narrow neck with a dilated horse shoe shaped section of 

 the vas deferens, which lies in the segment in front of that which con- 

 tains the funnel; the vas deferens then again narrows and a second 

 time perforates the intersegmental septum. The two vasa deferentia 

 of each side retain their distinctness, and open 'into about the middle 

 of a large glandular atrium which is tubular in form and much like 

 that of Acanthodrilus ; its minute structure is also similar to that of 

 Acanthodrilus. Neither the vasa deferentia nor the atria are furnished 

 with the thick muscular coat which is so prominent a feature of these 

 organs in Eudrilus (see Beddard j)Contributions to the Anatomy of 

 Earthworms«, Proc. Zool. Soc. 1887. pi. XXXIII figs. 8, 9, 10, 13, 16). 

 There is at most a trace of the musculature of the atrium, which how- 

 ever, as in other Earthworms, becomes thickened upon the proximal 

 part of the tube which is connected with the external aperture. The 

 two atria open by a single orifice situated at the apex of a muscular 

 penis which is placed on the border line between segments XVII. 

 —XVIII. 



The ovaries are paired and lie in segment XIII.; each is en- 

 closed in a separate coelomic sac, which contains a por- 

 tion of the nephridium belonging to its segment; the two ovarian 

 sacs communicate with each other by a narrow tube like sac, and also 

 with a large sac, which forms a complete ring encircling the oesophagus 

 and is continued into an extensive sac passing along the dorsal surface 



