601 



nerve does nut come into connection with the Gasserian ganglion in 

 Urodela . 



The important point here established is that the Fasciculus com- 

 munis enters the Palatine branch of the Facial, a nerve to the roof of 

 the mouth. According to Stirling (12), this branch is composed of 

 fibres to the mucous epithelium, to glands and to blood-vessels. Elim- 

 inating the fibres of the Sympathetic, it would be afferent and it follows 

 that the Fasciculus communis tract is, in all probability, afferent. 

 This view is further supported by its internal position in the dorsal 

 part of the medulla and by the fact that it constitutes the bulk of the 

 Glossopharyngeal . 



The other, coarse-fibred, division of the ventral VII. is the Hyo- 

 mandibular. Although it is for a short distance imbedded in the outer 

 ventral part of the Gasserian ganglion, it retains its continuity through- 

 out. In the Urodele forms examined and in those described by 

 Fischer (5), neither this nerve nore the Palatine comes into contact 

 with the Gasserian ganglion. Its position in Anura is not improbably 

 due to the greater development of the Auditory capsule, which pushes 

 it forward. As the Hyomandibular leaves the Gasserian ganglion, it 

 receives a number of the fine Palatine fibres. This is probably also the 

 case in Urodele forms and is likewise inferred by E wart as probable 

 in Elasmobranchs from macroscopic appearances (4). 



Thus the Hyomandibular consists of three components: one from 

 the ventral root of the ventral VII. (VII. ab] ^ coarse-fibred, one from 

 the dorsal VII. (VII. b), coarse-fibred and sensory, and one ultimately 

 from the Fasciculus communis (VII. aa), fine-fibred and sensory. Un- 

 fortunately the ultimate distribution of these parts is complicated by 

 the fact that the Hyomandibular receives a communicating branch 

 from the IX. -{- X. group. It would seem highly probable, however, 

 that the Kamus mandibularis, which is fine-fibred and supplies parts of 

 the oral cavity, derives its fibres from the Fasciculus communis com- 

 ponent (VII. a a). The dorsal VII. (VII. b) component must be regarded 

 (see below) as supplying cutaneous branches. From the joint nerve 

 formed by the Hyomandibular and the communicating branch from 

 the IX. -f- X. are also given off a number of motor branches. I believe 

 these are, in part at least, supplied by the remaining component, i. e. 

 VII. ab, especially as this has the position and characteristics of a 

 motor root. 



3. The Nerves of the Lateral Sense Organs. 



b) The dorsal VII. (»V.— VII. urh of O shorn). This is given off 

 from the dorsal part of the medulla immediately above the Auditory 



