and lifting of gill nets and other gear when 

 boats were not available. 



We used several types of equipment to col- 

 lect fishes. Beach seines and gill nets of 

 various lengths and mesh sizes, tow nets 

 3.3 m. and 1 m. in diameter, small otter 

 trawls, and small. meshed trap nets and fyke 

 nets were used routinely through most of the 

 Naknek system. Much of this equipment was 

 also fished in other areas, although trawls 

 and tow nets required powerboats, available 

 only on the larger Naknek system lakes. Other 

 sampling equipment included fish toxicants, 

 sport fishing gear, hand nets, fish spears, 

 small bobbinet "habitat" seines, and small- 

 meshed collapsible nylon traps. Details on 

 the use of much of this equipment have been 

 discussed elsewhere (Hartman, Heard, and 

 Strickland,'' Ellis,^ Heard, 1962; Merrell, 

 1964). 



Fish weirs or counting fences operated by 

 personnel of the Bureau of Commercial Fish- 

 eries at various times in the Naknek systenn 

 on Naknek River, Brooks River, American 

 Creek, Hardscrabble Creek, Hidden Creek, 

 and West Creek, and along Shelikof Strait at 

 the outlet of lower Kaflia Bay Lake have all 



contributed information on the occurrence and 

 distribution of fishes considered in this report. 



Fish collections were processed by identi- 

 fying and counting the number of each species 

 collected. In some collections, length and 

 weights were recorded, and scales were col- 

 lected for age analysis. Lengths of fish, except 

 where otherwise noted, are reported as fork 

 length. Field records from all Shelikof Strait 

 and King Salmon River system collections, 

 plus many of the Naknek system collections, 

 are filed in the museum of the Bureau of Com- 

 mercial Fisheries Biological Laboratory at 

 Auke Bay, Alaska. 



Identifications of fishes were made by the 

 authors except where otherwise referenced. 

 The principal taxonomic keys used were by 

 Wilimovsky' and Clemens and Wilby (1961). 

 Meristic counts follow Hubbs and Lagler 

 (1958). Representative samples of all species 

 collected in each of the principal drainages of 

 the monument have been deposited either in 

 the museum of the Laboratory at Auke Bay or 

 in the Oregon State University fish collection. 

 Nomenclature of fishes follows the American 

 Fisheries Society (I960) except where other- 

 wise noted. 



ANNOTATED LIST OF SPECIES 



The data on occurrence and distribution of 

 fresh-water fishes in Katmai National Monu- 

 ment are presented in the following annotated 

 list of 24 species from nine families known to 

 occur in the monument. Within the family 

 groups, six families --Petromyzontidae, Os- 

 meridae, Umbridae, Esocidae, Catostomidae, 

 and Gadidae--have only one species; whereas 

 two families--Gasterosteidae and Cottidae-- 

 have two species each; and one large family-- 

 Salmonidae- -has 14 species. In addition to 

 infornnation on fresh-water forms, we provide 

 data on four euryhaline species, each from 

 a separate family, that occur in or near fresh 

 waters of the monument. Besides data on the 

 occurrence and distribution of each species, 

 the annotations include information on the life 

 history and biology from various parts of the 

 monument. Such information generally is not 

 available from other sources. The quantity of 

 this information varies greatly from species 

 to species--a result of circumstance, not 

 design. 



Hartman, Wilbur L., William R. Heard, and Charles 

 W. Strickland. 1962. Red salmon studies at Brooks Lake 

 Biological Field Station, 1961. Manuscript on file, Bur. 

 Commer. Fish. Biol. Lab., Auke Bay, Alaska, 53 pp. 



^ Ellis, Robert J. 1963. The abundance and distribution 

 of juvenile red salmon and associated species in lakes of 

 the Naknek River system and Karluk Lake. Manuscript on 

 file. Bur. Commer. Fish. Biol. Lab., Auke Bay, Alaska, 

 80 pp. 



FRESH-WATER FORMS 



Many of the fishes occurring in fresh-water 

 areas of Katmai National Monument either are 

 or can be anadromous. A few forms that ap- 

 parently are confined to fresh water through- 

 out their life in southwest Alaska occur in 

 estuarine environments in the Arctic (Walters, 

 1955). Only three families in the monument-- 

 Umbridae, Esocidae, and Catostomidae--are 

 considered by Miller (1958) to be primary 

 fresh-water fishes. All 24 forms listed below, 

 however, at least spawn and spend their juve- 

 nile stages in fresh waters of the monument. 



Petromyzontidae 



Arctic lamprey, Lampetra japonica (Mar- 

 tens).-- Arctic lannprey were collected only in 

 the Naknek system, where fresh-water and 

 anadromous forms coexist. We collected ma- 

 ture, spawning, or spent individuals of the 

 smaller fresh-water form in May and June 

 fronn Coville, Grosvenor, and Brooks Rivers; 

 from most tributaries of Brooks Lake; from 

 two small tributaries of Naknek Lake, one in 

 North Arm and one on the south shore of the 

 lake near the monument boundary; and from 

 a tributary of Coville Lake about 1.6 km. 

 north of the lake outlet. Only one specimen of 



Wilimovsky, Norman J. 1958. Provisional keys to the 

 fishes of Alaska. Manuscript on file. Bur. Commer. Fish. 

 Biol. Lab., Auke Bay, Alaska, 113 pp. 



