\ 



Dr. W. H. Thomas, Assistant Research Biolo- 

 gist, July 1957-June 1960. 



Mr. R. C. Griffiths, Graduate Research Biolo- 

 gist, July 1959-June 1960. 



Mr, G. T. Barlow, Senior Electronics Techni- 

 cian (half-time), July 1958-June I960. 



Mr. R. J. Linn, Senior Marine Technician, 

 July 1957-June 1960. 



Mr. A. D. Reith, Senior Marine Technician, 

 March 1958-June 1960. 



Mr. M. L. Young, Electronics Technician, 

 December 1957-February 1959. 



Mr. R. V. Fidler, Laboratory Technician, April 

 1959-June 1960. 



Mrs. M. A. Miller, Senior Draftsman (half- 

 time), July 1958-June 1959. 



Mrs. J. F. Scotten, Secretary-Stenographer, 

 October 1957-June 1960. 



Mrs. J. L. Connell, Engineering Aid, November 

 1958-June I960. 



The following investigators were supported 

 from STOR funds in other ways: 



Prof. W. H. Munk 

 Prof. J. D. Isaacs 

 Prof. C. S. Cox 



Mr, J. M. Snodgrass 

 Dr. J. A. Knauss 

 Mr. G. 1. Roden 



Some of the above-mentioned persons v/ere 

 supported in part from other contract funds, 

 especially from the Office of Naval Research 

 and the Marine Research Committee of the 

 State of California, as indicated specifically 

 elsewhere in the paper. 



The Scripps Institution provided a group of 

 advisers called the Tuna Oceanography Ad- 

 visory Panel of the Research Advisory Council, 

 which had 12 meetings. Its membership on 

 June 30, I960, was as follows: 



Dr. E. H. Ahlstrom (Bureau of Commercial 



Fisheries) 

 Prof. R, S. Arthur 

 Dr. M. Blackburn, Chairman 

 Prof. C. L. Hubbs 

 Prof. J. D. Isaacs 

 Mr. J. L. Reid 

 Dr. M. B. Schaefer (Inter- American Tropical 



Tuna Commission) 

 Prof. C. D. Wheelock 

 Dr. W. S. Wooster 



The Bureau of Commercial Fisheries ad- 

 vised the Program Director from time to time 

 through John C. Marr, formerly Director of 

 the Bureau's Biological Laboratory at La Jolla, 

 and subsequently through Gerald V, Howard, 

 Director of the Biological Laboratory at San 

 Diego. There were frequent meetings with 

 these and other representatives of the Bureau. 



members of the Advisory Panel and gave 

 valuable counsel in other ways. 



There was close cooperation between STOR 

 and the Inter-American Tropical Tuna Com- 

 mission. The Commission assisted in many 

 ways, especially by providing data on tuna 

 catch per unit fishing effort and by helping to 

 staff cruises. 



The Bureau of Commercial Fisheries Bio- 

 logical Laboratories at La Jolla, San Diego, 

 Stanford, and Honolulu assisted by providing in- 

 formation, and in other practical ways. 



SCOPE AND PLAN OF THE INVESTIGATION 



Figure 1 shows the main parts of the in- 

 vestigation by subject and area. It does not 

 show the whole region of the fishery, which 

 extends broadly from latitude 30 N. to 20 S. 

 and up to about 700 miles from the mainland 

 coast, or the whole region of the investiga- 

 tion, which included the waters off Peru. 



The relationships of the items in figure 1, 

 to each other and to the main purpose of the 

 investigation, may be understood by reference 

 to table 1, which shows schematically how the 

 work was planned and carried out. 



In explaining table 1 it is convenient to start 

 at the bottom (Parts of energy chain studied). 

 It was assumed that atmospheric conditions 

 have effects on physical and chemical condi- 

 tions in the upper ocean which in turn affect 

 the availability of small biota (phytoplankton, 

 zooplankton, and micronekton, all in the tuna 

 food chain) and therefore the availability of 

 tuna (see Introduction). Attempts to identify 

 and study these effects, together with the 

 associated methodological work, are discussed 

 in the following sections of the paper: 



Statistical analysis of ocean-atmosphere 

 relationships 



Physical features and processes in the ocean 



Methods in physical oceanography: evalua- 

 tion and development 



Light, nutrients, and biota: methods and 

 experiments 



Light, nutrients, and biota: statistical 

 analysis of ocean data 



Tuna ecology (in part). 



It was also considered that physico-chemical 

 conditions of the ocean might affect availability 

 of tuna directly, as well as indirectly through 

 the small biota. This subject is dealt with in 

 the following sections of the paper: 



Recent changes in the ocean 

 Tuna ecology (in part). 



The late Townsend Cromwell and Dr. Bell 

 M. Shimada, of the Tuna Comnnission, were 



It was seen that such researches would have 

 their greatest practical value to the fishery if 



