Manchester Alenioirs, Vol. liv. (1910), No. 17. 5 



where it parts company with it. On the other hand we 

 have the very strong evidence already detailed that this 

 point is not morphologically a true node, and further, the 

 absence of any indication that the main protoxylem-mass 

 was here connected with the sporangiophore-trace. I 

 have in no case been able to observe a true fusion of the 

 trace with the stelar bundle, while, on the contrary, in 

 one section belonging to Mr. Watson (A90) the trace can 

 be followed quite uninterruptedly almost half-way down 

 to the subjacent node. In some cases the traces may be 

 distinguished in transverse sections below the sporangio- 

 phores as more or less projecting masses of tracheids on 

 either side of the main bundles (see Photo. 3 of Plate), 

 reminding one of the condition of things in Palcso- 

 stachya. As in the latter cone also I have observed in 

 one or two instances what I believe to represent small 

 independent carinal canals belonging to the traces 

 (see Photo. 5 of Plate). In the great majority of cases, 

 however, any tracheids which may have separated these 

 canals from that belonging to the main bundle have 

 broken down, the result being a single tangentially 

 elongated canal, often broadly V-shaped in transverse 

 section. Indeed, the whole trace is commonly broken 

 down, leaving a space on either side of the mass of 

 tracheids representing the stelar bundle. 



With a view to further investigating, among other 

 things, the relation of these traces, I obtained from Mr. 

 Lomax a block containing Calaniostachys and prepared 

 serial sections by the method already mentioned. One 

 part of this series shows very clearly the relations of the 

 traces in the manner just described, as represented in the 

 annexed sketches {Fig. 2, rt, b, c). The section is oblique, 

 and tangential to the bundle. The trace on the left has de- 

 generated and is represented by a space ; on the right, the 



