12 HiCKLING, Anatomy of CalaniostacJiys Binncyana. 



case in C. Binntyana. Renault states that the same was 

 the case in his cone of Annularia longifolia (Renault '73). 

 It was evidently so in Calajnostachys Ludwigii,^r\d Weiss 

 states that in impressions of C. sjtperba the ribs on the 

 axis do not alternate at the nodes. His figures show the 

 same condition in Stachanjiularia tiibercidata. The only 

 doubtful cases seem to be Renault's cones described in 

 his " Flore Fossile" ; regarding these neither the text nor 

 the illustrations are very satisfactory on this point, but as 

 a whole the evidence seems to favour superposition. 



If the contentions in this paper are correct, we may 

 say that it is a further general characteristic of calamitean 

 cones that the vascular supply of the sporangiophores 

 arises from the same node which supplies the whorl of 

 bracts below. This is the case in PalcBostacJiya, Renault 

 shows it to be the case in Bruckuiaimia {Arthropifostac/iys) 

 Decaisvei and B. Grand' Euryi, as well as in Calaniodendro- 

 stac/iys Ze27/er2 (Renault 'g6), and 1 have endeavoured to 

 show that it is the case also in CalaniostacJiys Binneyana. 

 Hence it holds for all the principle types of calamitean 

 cone. 



The point of insertion of the sporangiophores in the 

 internode is obviously very variable. In the PalcEostacJiya 

 type they are adherent to the bracts below, in the Stach- 

 annularia t\'pe to the whorl above, and in Calamnstachys 

 more or less in the middle. 



The number of appendages in a whorl, and the 

 number of bundles in the stele are likewise very variable 

 features. All that can be said is that multiples of three 

 predominate. 



There are two points of morphological importance in 

 which CalaniostacJiys Binneyana and its close ally C. 

 Lndivigii are contrasted with the other cones whose 

 structure is known (except, of course, C. Caslicana and 



