1913] YAMANOUCHI—ZANARDINIA 15 
gametes. In early prophase, the nucleus shows 44 chromosomes, all 
alike both in size and in shape (fig. 35). In middle prophase the 
chromosomes become more compact (figs. 36, 37). During the for- 
mation of the spindle, the nuclear membrane disappears and the 
equatorial plate is established (figs. 38, 39). The polar view of the 
plate shows 44 chromosomes (figs. 40,41). Each chromosome splits 
longitudinally and half of each proceeds to each pole (figs. 42, 43). 
The growth of the sporelings holds no strict relation to the mitosis 
within; the mitosis may take place before the sporeling begins to 
elongate (figs. 17, 39, 
gates while the nucle- 
us is in the resting 
condition (figs. 17, 34). 
The axis of the first 
division, as a rule, is 
perpendicular to the 
growing axis of the 
sporeling. After telo- 
phase the sporeling 
is divided into two 
cells (fig. 44). The 
nuclei in the two cells 
divide either stmul- Fic, 13.—Female gametangia with 4 tiers of cells: 
taneously (fig. 49) or a and b, a nucleus in each with chromatin accumula- 
one after the other _ tions around the outside of da nucleus 
(figs. 4 5~48). The in each in prophase, showing 22 chromosomes; ¢; 
nucleus in anaphase viewed from the pole, showing 22 
chromosomes. 


number of chromo- 
Somes appearing at 
prophase (fig. 44) and metaphase (fig. 45) is 44. After the 3-celled 
Stage, the growth of the sporeling is not very uniform. The 
numerous cases observed showed great variability in the method of 
developing into the multicellular condition. But one principal 
fact that holds true in almost all cases is that when the sporeling 
has reached the 2 or 3-celled stage, the cell divisions occur chiefly 
in a single terminal cell or in two upper cells, while the basal cell 
