136 BOTANICAL GAZETTE [AUGUST - 
funiculus is laterally attached. There are the usual two integu- 
ments, which grow well out beyond the tip of the nucellus (fig. 33). 
The ovule first appears as a lateral projection nearly filling the 
cavity of the immature carpel. As the latter enlarges, the ovule 
keeps pace with it. The integuments begin to appear before it 
assumes the anatropous position (fig. 34). The primary arche- 
sporial cell is differentiated about simultaneously with the appear- 
ance of the integument, and even in its resting condition may be 
recognized by its greater size (fig. 35). After, or simultaneously 
with, its differentiation, the tip cells of the nucellus divide by 
periclinal walls, forming two cell layers outside the archesporium. 
No primary parietal cell is formed. The primary archesporial 
cell is in this species the spore mother cell, and develops directly 
into an embryo sac without the previous formation of a row of 
megaspores. The first division occurs early (fig. 36). The later 
divisions follow in the usual order (figs. 37-39), and before their 
completion the integuments have reached complete maturity, 
closing over the tip of the nucellus. 
The account just given represents the normal development of 
the ovule and embryo sac in Richardia and holds true in perhaps 
5 Per cent of the plants examined. In most cases, however, the 
plants not only fail to set their seed, but will not do so even when 
artificially pollinated. I refer here solely to the plant as I have 
observed it growing in local greenhouses, and to the individuals 
(some 150) on which I have tried the experiment of artificial pollina- 
tion. The observation of gardeners, so far as they have been con- 
sulted, confirms the conclusion that the vast majority of plants 
of the strains here cultivated, and under the conditions existing 
in local greenhouses and gardens, cannot be made to produce seed. 
BURBANK, on the other hand, reports that the Richardia as it 
8tows wild in California matures seed in great abundance.’ 
The behavior of the artificially pollinated ovaries is peculiar. 
Ovaries not artificially pollinated shrivel in 10-15 days after 
teaching maturity. Pollinated ovaries behave otherwise. A large 
sas xcoues lily, in all its varieties, blooms and seeds in the utmost pes 
, a, setting big plump seeds in abundance. This includes all the yellow, 
white, and black varieties, J think the cause of not setting seeds in the east is wholly 
cli ics + 
limatic,”—From a private letter addressed to the writer. 
