310 BOTANICAL GAZETTE [OCTOBER 
tudinally wrinkled in the internodes above. The epidermis is thin- 
walled, and shows two types of hairs, pointed and glandular. The 
pointed hairs consist of a few cells in one row, covered with a 
granular cuticle; while the glandular hairs have a two-celled head 
borne on a single foot-cell, or on a long stalk of several cells in a 
single row; the sessile glandular hairs are especially frequent on the 
basal internodes, while the long-stalked hairs abound in the 
inflorescence. The pointed hairs are frequently curved to almost 
hooked, and occur especially on the upper internodes. Within the 
epidermis are a few (one or two) continuous strata of collenchyma, 
but only in the aerial internodes, not in the subterranean. The 
cortical parenchyma attains its greatest width in the short inter- 
nodes of the pseudo-rhizome, where it consists of about 20 thin- 
walled compact layers; no crystals and no deposits of starch were 
observed in any part of the cortex. 
The endodermis is present throughout the stem, but it is not 
always distinct, since the Casparyan spots are sometimes difficult 
to detect; besides, the individual cells are uniformly thin-walled, 
and of the same shape and lumen as the adjoining cortical paren- 
chyma; moreover, the endodermis does not contain starch. 
Bordering on the inner face of the endodermis is the so-called 
pericycle, relatively poorly developed in Phryma; as a closed sheath 
of stereids this tissue occurs only in the inflorescence, when the 
fruits have matured; in the pseudo-rhizome it is represented merely 
by parenchyma with isolated strands of stereome; while in the 
other internodes it is either uniformly thin-walled or interspersed 
with a few stereids, but with no regularity. 
The herbaceous stem of Phryma does not increase much in 
thickness, and beyond the formation of secondary mestome strands 
no other secondary tissues were observed. The primary mestome 
strands are thus readily visible in all the internodes, and they are 
strictly collateral; there are 6 in the cylindric epicotyl of the seed- 
ling, and they are arranged in two broad groups; in the mature 
plant the mestome strands constitute 4 broad strands, one in each 
angle of the quadrangular stele. These primary strands contain 
leptome, cambium, and short rays of hadrome, in which the young 
vessels, reticulated and scalariform, are much wider than the 
