316 BOTANICAL GAZETTE [OCTOBER 
from Nippon elliptic, while the lower ones were ovate-oblong; all 
the leaves of a specimen from Kansas were lanceolate; while all the 
leaves of a plant from Georgia were ovate. The leaf margin was 
observed to vary to the same extent from crenate to serrate, even 
on the same individual. It is thus somewhat difficult to define the 
typical leaf-form in Phryma; it varies much, but the ovate outline 
may be the fundamental one. 
The internal structure is bifacial, the stomata being confined to 
the dorsal face, and the chlorenchyma being differentiated into a 
ventral palisade tissue and a dorsal pneumatic tissue. On both 
faces of the blade the cuticle is thin and smooth except above and 
below the veins, where it is wrinkled. The epidermis is a little 
thick-walled where it covers the veins, and the cell-lumen is some- 
what wider on the ventral face than on the dorsal (fig. 17). Viewed 
in superficial sections, the lateral cell walls of the epidermis are 
undulate on both faces of the blade, and the stomata lack sub- 
‘sidiary cells. Short, almost sessile, glandular hairs (figs. 12, 13) 
abound on both faces of the leaf, and, as may be seen from the draw- 
ings, the head is two-celled. Besides these glandular hairs there 
are also, and especially on the ventral face, some long, pointed, 
multicellular hairs (in one row) with cuticular pearls. The chloren- 
chyma consists of a single layer of very short and plump palisade 
cells (P in fig. 17) covering three strata of an open pneumatic 
‘tissué (P* in fig. 17): in superficial sections this pneumatic tissue 
shows intercellular spaces of considerable width (fig. 18). 
The mechanical tissues are poorly represented, there being only 
a few hypodermal layers of collenchyma above and below the mid- 
rib and the secondaries, and also the leptome of the midrib is sup- 
ported by an arch of thin-walled stereome (S in fig. 19). A broad, 
thin-walled water-storage tissue surrounds the midrib, but there 
1$ no endodermis; the lateral veins, on the other hand, are sur 
rounded by green parenchyma sheaths (fig. 17). All the mestome 
strands are collateral, and the median is the broadest. The 
petiole, when examined just beneath the blade, shows exactly the 
same structure as the midrib, except that it contains two very thin 
mestome strands, one on each side of the median. F inally, it may 
be mentioned that the throat of the corolla of the flower is not 

