108 BOTANICAL GAZETTE [FEBRUARY 
He finds that the archegonium in liverworts exhibits both inter- 
calary and terminal growth; that in mosses this terminal growth 
contributes notably to the length of the organ, five or six adven- 
titious segments being formed at the expense of the apical cell; 
that the terminal cell does not contribute at all to the canal 
cells, either in mosses or liverworts; that the neck canal cells 
all arise from an initial cut from the mother cell of the egg; 
that two ventral canal cells are occasionally found in Sphaero- 
carpus, and one such case was noted in Marchantia; and that the 
ventral canal cell in Marchantia is capable in rare cases of being 
fertilized in place of the egg. 
Goebel (18), having examined the development of archegonia 
in Mnium undulatum, is unable to sustain Gayet’s views, but cor- 
roborates those of Kiihn, Janczewski, and Campbell, namely that 
the moss archegonium develops in part by segments (some of 
which are axial) cut from an apical cell; and in this regard is to 
be distinguished from the liverwort archegonium with its inactive 
cover cell. 
EARLY STAGES OF THE ARCHEGONIUM., 
In early stages archegonia and antheridia cannot be dis- 
tinguished. Enlarged papilliform cells arise from the surface of 
the receptacle, and segments are cut off right and left from a two- 
sided apical cell (figs. z-r0). This method, just as characteristic 
for archegonia as for antheridia, agrees with Hofmeister’s account 
(1), but differs from the accounts of Leitgeb (4), Kibn (8), 
Janczewski (9), Campbell (13), Gayet (15), and Goebel (18) in 
the absence of a wall that cuts off a basal cell from the arche- 
gonium initial or the archegonium mother-cell. When six t 
eight segments have been cut off, the two-sided apical cell is 
transformed into a three-sided one, a process that introduces @ 
history distinctly characteristic of the archegonium. 
The first indication of this change that marks the organ as an 
archegonium is seen in a division of the apical cell which results 
in the introduction of a curved wall more nearly vertical than 
that which cuts off any preceding segment. The approximately 
vertical position of this wall causes it to fall upon the same wa 
below as that which the wall immediately preceding joins (AES: 
