110 BOTANICAL GAZETTE [ FEBRUARY 
THE TERMINAL CELL AND THE CANAL CELLS, 
The most difficult part of the research is to determine the 
behavior of the terminal cell and that of the canal cells from 
this period. That the origin of the axial row, including the 
points just mentioned, is a difficult problem is evident from the 
conflicting accounts of those who have attempted to solve it. 
There is quite general agreement, however, that the terminal cell 
in the mosses does not cease activity after cutting off the primary 
peripheral cells and the inner cell, as does the cover cell in the 
liverworts, but continues to increase the length of the arche- 
gonium neck by cutting successive peripheral segments from its 
three lateral faces. 
The status of the problem of the origin of the canal series 
may be summarized as follows: Hofmeister’s (1) idea that the 
canal series arises from one of the four original pedicel rows as 
triangular tangential segments of its cells is mentioned only as a 
matter of history, the erroneous conception soon being aban- 
doned; Kuhn (8, p. 31) claims that all the cells of the axial row 
are cut from the base of the apical cell (cf. his diagram, p/. 70, 
Sig. 71); Janczewski (9, pp. 412, 413) says that the numerous 
canal cells are of diverse origins, the lower arising through trans- 
verse divisions of segments cut from the base of the apical cell; 
Campbell (13, p. 19) says that the segments cut from the base 
of the apical cell constitute the axial row of neck canal cells, 
which, so far as could be determined, do not divide after they 
are first formed; according to Gayet (15, p. 241), all the neck 
canal cells have the same origin, all arising from an initial cut 
from the mother-cell of the oosphere, and no segments being cut 
from the base of the terminal cell; and finally Goebel (18, p. 
244) holds that the moss archegonium is distinguished from that 
of the liverwort by its characteristic growth from an apical cell, 
which also contributes canal cells, fi 
of twenty canal cells, 
Three sources are thus alleged for the canal cells: (1) the 
apical cell with intercalary additions; (2) the apical cell alone; 
(3) not the apical cell at all, but one at the opposite end of the 
row. Gayet alone holds the process to be like that in the liver- 
guring such a cell in a row 
