114 BOTANICAL GAZETTE [ FEBRUARY 
drawn: (1) The terminal cell is triangular in cross-section, 
inclosed by three peripheral cells which have been cut from it 
successively (figs. 48,6, 49,2). No mitotic figures were found, 
but the position of the walls and nuclei indicate their origin, 
(2) The three peripheral cells divide longitudinally and produce a 
neck of seven rows—six peripheral and one central (fig. 48, ¢, d). 
These divisions are not simultaneous (fig. 49, 2) and appear early, 
for the neck has six peripheral cells at the level of the second 
nucleus, less than 0.5™" from the apex (fig. 48,¢). (3) The neck 
has two peripheral layers near the venter (fig. 48,7). (4) The 
canal is hexagonal in outline and maintains this character till the 
egg is formed (jig. 48,7), and the venter wall breaks up into 
many cells (fig. 49, g). (5) Certain large cells, lying just below 
the egg and having the appearance in longitudinal sections of 
being members of the row (figs. 26, 27), are found here also, but 
are clearly no part of the row (fig. 48, #,/). (6) The pedicel 
bears throughout its length the quadrant character given to it 
by the early longitudinal divisions of the pedicel segments. 
Beginning at the base, the development upward is through quad- 
rant, octant, and intermediate stages to the base of the archego- 
nium proper, where the pedicel is twelve-celled in cross-section. 
At this point each quadrant consists of a center cell and two 
peripheral cells (fig. 49, g—h). 
THE PERIPHERAL CELLS. 
From the mode of origin of the neck of an archegonium of 
six rows, as already described (fig. 48, c, d), itis evident that the 
cells forming any row do not stand directly over one another, 
but alternate spirally around it, since they are cut off succes- 
sively around the apex (cf Janczewski, 9, p. 409). The longi- 
tudinal divisions which produce the six neck cells are usually 
followed by two or more transverse divisions in each cell, giving 
rise to vertical groups of four’ (fig. 23). In the early pedicel 
segments only one division occurs, and in some of the later 
segments division occurs in only one of the two cells arising from 
the first division, giving rise in such cases to groups of only two 
or three cells. The axes of these groups come later to stand in 
an oblique direction (fig. 23). 
