122 BOTANICAL GAZETTE [FEBRUARY 
5. Longitudinal radial divisions follow in these segments and 
form a pedicel of four rows. 
6. In the young archegonium the two-sided apical cell gives 
place to a three-sided one which is truncate. 
7. This terminal cell divides transversely soon after its forma- 
tion, giving rise to the first cell of the axial row. 
8. The terminal cell adds to the growth of the neck by seg- 
ments cut from its three lateral faces, and to the growth of the 
axial row by segments cut from its truncate face. 
9. Growth in length of the archegonium neck is intercalary 
as well as apical in both the neck and canal rows. 
10. The canal series is not always a single straight row, but 
sometimes double for a greater or less distance. 
11. The peripheral cells are not in vertical rows, but alternate 
with preceding cells; and torsion is a consequence of this feature. 
12. The egg and ventral canal cells are often of nearly equal 
size. 
13. The cells of a axial row are homologous with each other 
and with sperm mother-cells. 
14. Archegonia and antheridia are homologous structures 
throughout. 
15. The breaking up of the axial row into sperm mother-cells 
is an expression of partial atavism on the part of the archegonium. 
16. The mass of indefinite cells in the apex of the arche- 
gonium neck is the vestige of this atavistic Re oP oe): on the part 
of es archegonium. 
. Archegonia and antheridia probably had a common origin. 
Hyppotacteal ey the structure from which they have been derived 
was an asexual multilocular terminal zoosporangium of some 
primitive extinct member of the Chlorophyceae; the course for 
the archegonium being through a gametangium possessing more 
than one functioning gamete. 
HuLi BoTanicaL LagoraTory, 
cago. 
LITERATURE CITED. 
1. HOFMEISTER, W., Vergleichende Untersuchungen der Keimung, Entfal- 
tung und Fruchtbildung héherer Kryptogamen. Leipzig. 1851. 
