a 
ert 4 
3 
- the ridge has attained a little height it bifurcates, so that two 
1904 | LEWIS: ANOMALOUS DICOTYLEDONS 133 
have their origin as a very broad crescentic ridge of tissue. After 
lobes are formed. 
In older stages the two lobes are of equal size and the 
notches are of almost equal depth. fig. 23 represents the con- 
dition in the mature embryo. Section @ passes through the 
body of the embryo just below the growing point; 4 passes 
through the plumule; the remaining sections pass through the 
cotyledons. It will be observed that no cotyledonar tube is 
formed and that the edges of the cotyledons are not close 
together. 
The mature embryo is small, both hypocotyl and cotyledons 
| being short. The plumule has about the same development here 
as in the mature embryo of Podophyllum, being simply a small 
mound of tissue between the two cotyledons. The conducting 
tissue is beginning to develop as a central strand of narrow 
elongated cells, which divides into two branches, one going to 
each of the two cotyledons. Fig. 25 shows the shape of the 
mature embryo in longitudinal section. 
CAULOPHYLLUM THALICTROIDES. 
Material for the study of the younger stages in the develop- 
ment of the embryo of Caulophyllum was not. secured, but it is 
hoped that the investigation of the development of the embryo- 
sac and young embryo may soon be carried out. 
The youngest stages examined show the embryo at about the 
time when the cotyledons begin to develop. The embryo at this 
time shows a long, slender suspensor and a rather broad, short 
body (jig. 26). From this longitudinal section ie cor eecce 
seem to arise as two opposite ridges of about equal size, but it is 
impossible to say that this is the case, as cross-section series of 
this stage were not secured and series from embryos a little older 
do not support this view. A cross-section series from an embryo 
corresponding in size to the one represented by jig. 27 is shown 
in fig. 28. From this figure it is apparent that the cotyledons 
have a common base consisting of a low ridge of tissue which 
bifurcates to form two very symmetrical cotyledons. Older 
7 stages show that the bifurcations soon become of almost equal 
