1904 | SMITH: NUTRITION OF THE EGG IN ZAMIA 347 
the wall is formed of cellulose and contains a large number of 
canals showing genuine sieve plates, through which the proto- 
plasm of the cells of the endosperm layers communicates with 
the protoplasm of the egg.” 
Ikeno (5) in his paper on Cycas revoluta says: “Since Goro- 
schankin’s researches it is known that in the cycads the proto- 
plasm of the central cell and of the wall cells are in close con- 
nection by means of protoplasmic threads. It is easy to see that 
the material filling the nuclei of the wall cells, which in its natural 
condition is a half-fluid substance, flows out from there and goes 
to the central cell through these intercellular breaks. Indeed, I 
can find various stages of the transportation of this material to 
the central cell, showing as granulations in fixed material, where 
these granules lie just outside the nuclei, then in the protoplas- 
mic threads themselves, then in the central cell directly before 
the protoplasmic break. Also one finds a large mass of these 
granulations at the edge of the central cell, which have collected 
there, evidently coming completely from the neighboring wall 
Rélis:” 
Arnoldi claims that the nuclei of the jacket cells pass bodily 
into the egg through the large pores in the inner walls of the 
jacket cells. In writing of Pinus cembra he says (2): “I can show 
that I could observe out of a series of sections more than one 
hundred and fifty nuclei passing out of the jacket cells. It is 
also established through my observations that the small proteid 
vacuoles described by Goroschankin as Hofmeister’s nodules are 
only nuclei passing out of the jacket cells.” 
In writing on Cephalotaxus Fortunei he says (1): “In the first 
Stages of development of the jacket cells we find the httle 
_ fucleoli only half inside of the nuclei. In some later stages we 
find them also in the protoplasm of the jacket cells. Since these 
small granules are fully equal to those in the nuclei in all rela- 
tions of size, staining ability, etc., since they are first observed 
in the nucleus and later in the protoplasm, we may conclude that 
they pass from the nuclei into the protoplasm. In the proto- 
plasm of the jacket cells they are heaped up in great masses, 
and then pass over into the egg cell. The nodules appear in the 
