604 PROCEEDINGS OF THE ACADExVY OF [Oct., 



line; lateral margins of the sclerites narrowly dark brownish black; 

 sternites dark brown. Hypopygial sternites bright yellowish chest- 

 nut, tergites brown. 



HolotjTDe, cf, Nipigon, Algona District, Ontario, ,June 17, 191.3 

 (Dr. E. M. Walker). 



Allotype, 9 , topotypic. 



ParatjTDe, No. 1, cf , topotypic; No. 2, cf , type locality, June 18, 

 1913; No. 3, 9, North Fairhaven, Cayuga Co., N. Y., May 17, 

 1913, found dead in lake drift (Dr. J. G. Needham and Miss Emme- 

 line Moore). 



The type and paratype No. 1 is in the collection of the University 

 of Toronto. 



This insect is closest to P. replicata LinniEUs of Europe, but the 

 wings are darker colored, much more tinged with l^rowai; the vena- 

 tion, although similar in the persistence of the tip of i?i, shows a 

 tendency to the reduction of the radio-median cross-vein, the base 

 of cell 1st Mi more arcuated and other details. Grunberg's figure 

 of the male hypopygium of replicata^^ shows differences in the shape 

 of the 9th tergite and the conspicuous appendages of the 9th sternite. 

 The wing venation of the three knowni species of the genus are figured 

 on PI. XXV, replicata, fig. 9, neoxena, fig. 10, tipulina, fig. II. 



At this point it may be mentioned that there is a great difference 

 in the interpretation of the venation of the radial field of the wing 

 in this tribe of craneflies. Most authors have considered the 

 vein R: of the CylindrotominEe to represent a combined fusion of 

 A'. +2+3 from the tip of the wing backward. From a study of the 

 venation of the known species of this tribe, about a dozen in all, 

 it is seen that the above interpretation of a long backward fusion 

 of Ri+i+a is impossible and two other possible explanations are here 

 presented. Looking over the series of wings before me, it seems 

 that the vein hitherto considered as Rn-i+s is, in reality, Ri or /2j+, 

 alone, Ri becoming atrophied beyond the radial cros.s-vein rather 

 than obliterating this cro.s.s-vein and fusing with R,. This is proved 

 by the wings of Phalacrocera shown in the plate, in replicata and 

 neoxena, Ri being separated from Ri+s, whereas in tipulina the tip of 

 R, is atrophied beyond cross-vein r. A second possible interpretation 

 is that of considering the small cross-vein mentioned by Osten 

 Sacken as occurring in the costal cell beyond the tip of Sc and present 

 as a very indistinct vein ia many specimens (Liogma) as being the 



» Su>.s\va.sserfauna DcutsdihuKls, vol. 2.\, pt. 1, p. 33; 1910. 



