ILLUSTRATIONS OF INDIAN BOTANY. 213 



A one-celled ovary is one where the marghis of the carpellary leaves are not so far inflexed as 

 to meet and coalesce in the axis, and therefore leave an empty space in the centre ; a t\^'o- or more 

 celled one is where these margins meet and coalesce in the axis. " Of the first there are Uvo 

 degrees — first, where they meet in the periphery and are not inflexed, having the placentse either 

 central or parietal — second, where they are so far inflexed that they nearly meet but, owhig 

 to the edges becoming reflexed, do not cohere. In this last case the partition, in place of resem- 

 bling a room with a wall built across, resembles one divided by folding doors which, though they 

 may touch, can still be withdrawn without causing any breach of continuity. This last form 

 is not deemed a true partition, and ovaries in which it occurs are not considered truly 2-celled. 

 Such ovaries are said to be spuriously 2-celled, such is the ovary of Clcrodendron^ ViteXy 

 Premna^ and I believe some others of that tribe. For other examples of this form of ovary 

 see the figures of GentianecB^ OrohancacecB^ GesneriacecE^ &c. Such a structure seems never 

 to have been suspected in this order; hence Mr. Griffith's astonishment on finding it in the 

 genus Glossocarya^ which he describes as having an ''Ovarium 1-Ioculare ! placentse bilamellatiP, 

 lamellis recurvato-incurvis ; ovula 4 pendulo-apensis."" Schauer, however, either did not see the 

 fine of separation between the two semi-partitions or folding doors, or does not consider cohe- 

 sion necessary to the formation of a 2-celled ovary, and assigns to that genus that structure. 



Be that, however, as it may, I find a similar structure in other genera of the order, showing 

 an analogy if not a positive aflJnIty between Viticece and BignoniacecB^ and even Acaiithacece . 

 Symphoremece and Avicennece differ from both these forms in the structure of their ovary, 

 which is partially 2-celIed, that is, the partition wall, if I may so say, is not raised to the 

 ceihng, so that it is 2-celled below and one-celled above : which is my reason for wishing to 

 separate the former as a distinct and well marked tribe from Vitece. In Myoporace^^ an order 

 to which Amcennece is so nearly related that Brown referred it there, I do not find this structure. 

 In it, the ovules are really attached to the upper angles of the cells which are complete, at least 

 I find them so in the only species I have at hand to examine, namely, M. congestum and 

 acuminatum. This difference, though microscopic, seems one of considerable import, espe- 

 cially when combined, which I believe it is, with difference of habit. But on that point I 

 speak with diffidence, having so little ac(iuaintance with the order. Gmelina differs from the 

 Viteous genera above mentioned in having a distinctly four-celled ovary, with a single ovule 

 in each, pendulous from the upper central angle, just as in Myoporacece ; it also corresponds 

 with that order in the character of the anthers; and, as regards the corolla and fruit, I think 

 agrees even better than with Verbenacece. For these reasons I am disposed to view the genus 

 as the transition one, uniting the two orders and, perhaps, of the two more justly referable to 

 the former. Under this impression I have introduced a figure of Gmelina with the view of 



making known, througl 



Myoporacece 



approach each other, and have added in the upper corner of the plate some outhne sketches, 

 copied from Endlicher's Icones, of a species of that order which I think go far to prove that 

 Gmelina is erroneously placed in this order, as it only differs from the other m its 4-lobecI 



corolla. 



inflexed) 



Character of the Order. Flowers hermaphrodite, rarely polygamo-clioicous, 4-5-me- 

 rous, rarely more; regular or irregular or bilabiate, unlbracteate; bracts sometimes enlarging 

 after blooming. Calyx free, monosepalous, 4-5- rarely G-8-toothed persistent, more or less 

 enlarging with the fruit. Corolla hypogynous, monopetalous, tubular, deciduous ; limb 4-o- 

 sometimes G-12-lobed, usually unequal, secund or more or less perfectly bilabiate, rarely equal ; 

 „,„i!.-_.. . , . , ;. ri . •'_ i_ _ _ _„ „ i„i\^„^A\ ftfomens inserted on the tube ot the corolla, 



imous, and all fertile, or with a superior 

 J : cells generally opening longitudinally, 



'^Acvuiueriierous or ruaimentary uuc , aumcio ^-^^.^.^^ ■, ^^na 5^ ^ r j u«„„„ j +v,^ ^^iL 



parallel, divaricate or vertically superposed ; connectivum sometimes F«^ "^f ^^^^ «" V^' ^^^^^^^^^ 

 Ovary free, seated on an annular disk composed of 2 or 4 carpels; 2-4-celed with the margrns 

 of the carpels forming the primary partitions, or by these partitions, ^^^^^^^ VSl ^LT^Lt^^^ 

 the cavit/and introflexed, 4-8-celled. Ovules usually solitary m the cells Jj^^^b g^"^^^^^^^^^^ 

 collateral, or iwo opposite ; erect from the base, anatropous, in Verhenece, m the other tribes 



m 



e 



didyn 



