ON THE AWNS OF. NEPAUL BARLEY. 35 
its most rudimentary state. Its base lies towards the apex of the awn, 
whilst its axis is (in this example) so much inclined as to be nearly 
parallel to the axis of the awn. Between the cellular mass of which 
the bud is formed and the awn, is a thick rudimentary scale (6) 
which I shall name the “ basial leaf” of the bud; and it will be seen, 
that the mass of the bud adheres to it, except at its apex. This ** basial 
leaf ^ is invested with succulent hairs at the back and margin, and a 
few such hairs are also seen on the cellular mass of the bud itself. 
That the axis of this bud is not primarily or essentially so much 
inelined to the axis of the awn as in this instance, appears to be 
shown by the specimens figured at 9, 10, 11, where it has developed 
under the form of a sub-cylindrical column, but does not exhibit 
any very distinet traces of separation, or subdivision, into organized 
appendages of any kind. A transverse section of this columnar mass 
shows it to be composed of cellular tissue, within which green matter 
is deposited throughout the space running a short, distance within and 
concentric with the surface. In these instances the axis of the bud is 
perpendicular, or very nearly so, to the axis of the awn. But where 
the two axes are much inclined to each other, this has been owing (as 
it seems to me) to the base of the bud having been carried a little 
forward, subsequently to its development, by the extension of the 
wn 
The position of the apex of the basialleaf may here be observed 
marked out by a few hairs at the base of the column towards the 
side nearest the base of the awn. All the further modifications 
which I am about to describe appear to have resulted from the relative 
degree of influence that has been exerted between the developing forces 
respectively employed in extending the awn and the bud, wherever the 
axis of the latter has become more or less inclined to that of the former. 
If I may be allowed the use of the expression, “axis of development," 
to signify any axis whatever round which, and in relation to which, 
organized tissues have formed or are forming, we shall here have two 
such “axes of development” (that of the awn, and that of the bud) 
which become more or less inclined to each other, though normally at 
right angles. The results produced by the forces which collect and 
arrange the materials round these two axes respectively will therefore 
interfere, and we may expect to see a vast variety of external 
F2 d 
