316 DESCRIPTION OF SOME NEW PLANTS 
they are definite being those which are most nearly related to Bruni- 
acee, such as Tetracrypta and Trichocladus. Indeed, judging from the 
character of it given by Harvey, for I have no specimen to examine, 
ichocladus seems to exhibit more the habit of Bruniacee than Ha- 
mamelidacee, but it has the large embryo of the latter tribe.* 
The anthers in Bruniacee are introrse, and the cells burst by longitu- 
dinal slits. They are either ovate, with the cells diverging at the base, 
and attached by their back to the filament, or oblong, and attached by 
the base. In the character of the Order given in Lindley’s “ Vegetable 
Kingdom,” the anthers are distinctly stated to be turned outwards ; but 
in five genera which I have been able to examine, they are all turned 
inwards ; and such also are they represented in the whole of the genera 
figured by Brongniart in his memoir.t The filaments are either fili- 
form, subulate, or nearly wanting. 
As in all the other points of structure the anthers of Hamamelidacee 
are very variable. I have already stated that in the new genus Tetra- 
crypta they are not to be distinguished from those of Brunia. In 
Parrotia, Fothergilla, Bucklandia, and Sedgwickia, they are much the 
same as those of Ardouinia, Thamnia, and Tittmannia, being oblong, 
fixed by the base, and having the cells opening by longitudinal slits. 
In most of the other genera they are fixed by the base, but open by 
vertical valves, which in one instance, the Hamamelis Chinensis, Br., 
are deciduous. In all the other genera except Zustigma they are in- 
trorse. 
Tn connection with the stamens there is another point of saline 
between the two Orders, both having sterile stamens alternating with 
the fertile ones. In Brumiacee, however, they only occur in a 
single species of Brunia, B. nodiflora ; while in Hamamelidacee they 
are to be found in several of the genera. The existence of sterile 
stamens has not hitherto been distinctly stated to exist in Brunia, the 
petals, to which they adhere, being merely said to be bierested. That 
they are sterile stamens there can be no doubt, for an attached filament 
may be traced along the mesial line of the petal from the same point as 
that from which the fertile stamens take their origin, and are clearly 
seen to belong to the same series. ~The two crests which occupy the 
middle of the petal are abortive anthers. Thus there is a tendency in 
both tribes to produce as many stamens as there are sepals and petals. 
* “The genera of South African Plants," p. 127. + Annal. des sc. Août, 1826. 
