122 
were isolated, and the offspring of each were thus kept under 
observation. Much to my surprise one of the females pro- 
duced offspring of three types, the normal phase like the parent, 
a red phase with only the inner margin of the elytra black, and 
a black phase with a red cross-band behind the humeral angle 
of the elytra. JI experimented with these forms as much as 
the time at my disposal and the ill-effect of inbreeding would 
permit, but could discover no evidence of Mendelian inheri- 
tance except in the perfect segregation of the phases. A com- 
plete set of data obtained from these experiments with Cheilo- 
menes was submitted to a well-known student of Mendelian 
inheritance, Dr. John Detlefsen of the University of Illinois, 
and he was unwilling to venture any explanation of the 
phenomena presented, although loath to admit that the inheri- 
tance was not amenable to Mendelian laws. 
Having seen from my own experiments and from those of 
other workers, including Mr. A. F. Burgess, Professor R. A. 
Johnson, and Miss Miriam Palmer, that while the inheritance 
of ladybeetles is often segregative, its phenomena are not other- 
wise easily amenable to Mendelian interpretations, I was sur- 
prised and delighted to find a case of simple Mendelian in- 
heritance in the Australian ladybeetle, Coelophora inaequalis 
(Fabricius). This inheritance was coupled, moreover, with an 
example of a segregative but apparently non-Mendelian inheri- 
tance in an illuminating manner. 
Coelophora inaequalis, so well known to Hawaiian  ento- 
mologists, was an early introduction of Albert Koebele’s into 
these islands from Australia. As found here it exhibits a 
remarkable uniformity of markings, and shows no trace of the 
range of variation credited to it elsewhere? and, in fact, it 
represents the normal and most abundant of the three phases 
7 The geographical range of this species is reported to extend from 
Japan and the Philippines through the East Indian Islands to New 
Caledonia and Queensland, but I am convinced that the Philippine form 
really represents a similar but quite distinct species from the Australian. 
The fact that there are two species confused under this name only par- 
tially explains the reputed variability of inaequalis, as I have reason to 
believe that both have a similar range of variation, and that similar 
color phases are common to both. 
