IN NATURAL CLASSIFICATION. 593 
In the Iridee and Amaryllidee, the pollen is the same as 
in the Liliacee. 
In Scitaminee and its allies, the pollen is very pecu- 
liar, quite different from the foregoing. As far as I have 
ad an opportunity of examining it, it seemed spherical and 
hispid when dry, separated a diaphanous membrane from its 
surface when placed in water, and protruded three equi- 
distant, or club-shaped masses when placed in acids. 
In Orchidee the pollen consists of a variable number of 
vesicles cohering by their exines, sometimes reunited by a 
cellular growth into a club-shaped mass. 
Among the Achlamydeous and Monochlamydeous Dicoty- 
dons, the pollen is for the most part spherical or oval, 
opaque, and protruding a number of vesicles on its surface, 
caused by the swelling of the fovilla pushing out the intine, 
through pores of the exine. The Salicinee and Proteacee, 
however, have an elliptical transparent pollen, marked with 
a dark central line, which disappears when placed in water, 
a bubble of air being expelled, its figure at the same time 
becomes spherical, and it bursts at three equi-distant pores 
when put into acid, or at least becomes triangular by means 
of these re-agents. I have shown on a former occasion (vide 
Journal of Botany, Sept. 1840), that a triangular or three- 
lobed pollen sometimes occurs from the pollen having been 
developed under acid influence, as in Grevillea. 
That natural group composed of the Pedaliacee, Jasminee, 
Brunoniacee, Salvadoracee, Labiate, Verbenacee and Bora- 
ginee, whose common characters are, their being monopeta- 
lous dicotyledons, with superior ovaries, exalbuminose seeds, 
and erect ovules, have a pollen of the same type as that of the 
apetalous dicotyledons, namely, oval or spherical, opaque, 
and forming vesicles on the surface when placed in water. 
Boraginee, which differs from the others in having an ab- 
hilose radicle, is remarkable for its pollen, which appears to 
be formed, in the greater number of genera, of two vesicles 
contained in a common membrane, and bursting by two op- 
posite pores, at the point of juncture; in Borago, it is to be 
remarked the pollen is simple. 
