104 Macuieay Memorian Vobume. 
ganz verschiedenes Aussehen haben, die letzteren sind ausserordentlich viel volumi- 
nodser, sodann bilden die einen gar nicht die Fortsetzung der anderen, sondern sind 
ganz unabhiingig von ihnen. Ausserdem muss es aber auch auffallen, dass die 
kriiftigen Kaniile sich nicht weiter in die homogene ‘ Basalmembran’ verfolgen lassen.” 
To this I have to reply that the figure in question is not extremely diagrammatic 
—being in reality a very fair representation of what is to be seen in a section of the 
integument of 7. fasczata, and, as far as the cuticle, epidermis and basement 
membrane are concerned, a photograph would give a very similar picture. The 
parts of the canals that are situated in the epidermis are certainly wider than those 
in the cuticle ; in imperfectly preserved or partially macerated specimens they become 
very much more dilated. I did not make the lines in epidermis and cuticle coincide 
because, as a matter of fact, they do not do so. 
Let us proceed to examine Brandes’s own account of these layers. 
He states that he also finds in 7. dvevzcornzs an integument consisting of three 
layers. But he was not able with certainty to distinguish these from one another. 
The superficial part, he considers, might as well be regarded as a superficial part of . 
the whole “cuticular layer” which had undergone a certain modification through 
contact with the water, and the lowest layer—the basal membrane—may be reckoned 
as also part of the cuticle,* or as a delicate outer layer of the parenchyma, or as 
circular muscle-layer. But, he adds, the blame for these negative results may be laid 
on the smallness of the elements and their imperfect preservation. He then proceeds 
to describe in the middle layer the spaces above referred to but not previously noticed 
by me, and traces their connection with the pore-canals. 
All this is not readily to be reconciled with the passages I have quoted, and I 
am ata loss to see what is the ‘“falsche Deutung” which he regards himself as 
having detected. 
V.—Muscurar Layers. 
In the wall of the body there are three layers of muscular fibres—an external 
thinner layer with transversely or circularly arranged elements, an internal, much 
thicker layer in which the fibres run longitudinally, and an intermediate layer of 
intercrossing diagonal fibres. The first (Pl. x. figs. 1, 2, 7 and 8, c. m.) lies 
immediately below the basement membrane; in some of the species it is separated 
from the internal layer by a tolerably thick zone of parenchyma containing pigment. 
* T do not understand why Brandes should prefer to use the word cuticle here, seeing that he acknowledges the 
presence of nuclei. It is true that in the explanation of the plates he calls these “‘kernartige Gebilde”; but no uncertainty 
as to their nature is expressed in the text. To calla nucleated protoplasmic layer a cuticle is, to say the least, very 
unusual, 
