OF MOVEMENT IN PLANTS. 67 
may commence in a direction with the sun, or the reverse. Movement in either direction 
is by no means continued during the entire period of activity. Motion in one direction 
may soon be succeeded by movement in the other direction, one alternating with the 
other constantly. The dextrorse motion, for all the observations taken, aggregated 1622.10 
em., the sinistrorse amounted to 1400.95 cm., and the ratio of one to the other was, 
therefore, as 1 to 0.86. ; 
While this shows a tendency to equality of movement in the two directions—a ten- 
dency which might have been more pronounced had the observations embraced all the 
movements—an important relation bearing upon this point is to be observed between the 
latitudes and departures of movements. Also, the relation which these two directions of 
motion bear to one another must obviously be directly related to the location of the bands 
of more active tissue which induce the motion. The following table will exhibit the total 
latitudes and departures for all the tendrils :— 





Tendrils. Latitudes. Departures. Ratios. 
None ul 1244806 Ne dies | 249.70 | 1:2.01 
Cie eared aa ees HI Ca SEE 59251 | 116 60 | 1: 2.19 
rie APE) UMM sto San lfe se aaee | 269.40 | 1:2.10 
RARE men Rene etre 2100 RES 60.83 | 1 : 2.89 
RE Gares 3.37 ) 91.10 
285.17 489.65 | 1 :1.71 
CNE bec 241 80 J 398.55 J | 
CAPE RE heen TO ne | 266.55 | 1: 1.88 
| | | 
Te er 87 75 ) | 192.80 | 
| 135.50 } 262.10 | 1 : 1.93 
Et 47.75 | 69.30 | 
Wee Riie sed one 106.30] | 264.70 
| | 
Se ete 67.65 L| 227.15 | 203.03 || 612.93 | 1 : 2.69 
| | | | 
SE Ce 53.20 J | 144.50 J | 
| 
CT OU es tase ped See 2AM || osdoss 118.15 | 1: 0.95 
BALE = "| L 
Motals eee Ih gee. IGIGRERYA || Saeaaue 2445 21 
| | 1 : 2.04 
Means-:-c lle: | (LAS CENT 188 .09 
| 





An inspection of this table at once exhibits a most striking relation between latitudes 
and departures of motion. While in some cases there is a marked variation in the results, 
e.g., Nos. 4 and 9, yet these, as already seen, were tendrils which were only partially 
observed, and if we consider the mean result, which agrees with specific cases in which 
the entire action of the tendril was noted, we find the departures of motion to be just 
twice the latitudes. This indicates most conclusively, therefore, that the principal energy 
of circumnutation must be developed along the two sides of the tendril arm, and reference 
to our figures, as also the description of the histological elements, will at once show that 
it bears a most important relation to the three bands of vibrogen tissue. 
