198 CILIATE FIBRILLAR SYSTEMS 



ring surrounding (2) the basal granule of the cilium, and (3) the 

 "Nebenkorn" occurring at the right of the junction of the basal ring 

 and the interciliary fibril (p. 158). According to Klein (1931), the 

 elements of the basal apparatus include (1) a ring (Zirkularfibrille) 

 within which are usually three granules (Drierkorner). The central 

 granule is the basal granule, and the other two are Nebenkorner. The 

 discrepancy here is not only in the relative number of granules in the 

 basal apparatus but in their relationship, since von Gelei regards his 

 "Nebenkorn" as identical with Klein's basal granule, and accounts for 

 Klein's third granule as being only a thickening of the basal ring. 



Finally, mention may be made in this connection of an additional 

 system of fibrils described by Gabor von Gelei (1937) in three species 

 of Parameciujn, — P. caudatum, P. multhnkronucleata, and P. trichinum. 

 This third fibrillar complex, in addition to the two noted above, was 

 found at the level of and below the basal granules. Its fibrils spread 

 throughout the entire body surface, including the vestibule, where it 

 sends a thickened fibril into the cytopharynx between the membranelles. 



The general pattern of this complex, made up of longitudinal and 

 cross fibrils, resembled that of the outer fibrillar complex, the meshes 

 of the former being smaller and more numerous, however, than those 

 of the latter. Also, the course of the fibrils of this third complex were 

 more irregular. Its longitudinal fibrils become fewer in the middle region 

 of the body and at times are directed diagonally, even spirally. A splitting 

 of fibrils was occasionally observed, as well as variations in their thick- 

 ness. They may also anastomose and form a "Schaumgiter." 



The author could discover no connection between this fibrillar complex 

 and either of the other two. 



Apparently the most recent detailed account of the fibrillar system 

 of P. caudatum, including that of its cytostome, was made by Lund 

 (1933), working in Kofoid's laboratory. After comparing the descrip- 

 tions of earlier workers (Engelmann, 1880; Maupas, 1883; Schuberg, 

 1905; Rees, 1922; von Gelei, 1925-32; Klein, 1926-31; Jacobsen, 1931) 

 with his own findings, Lund concluded that previous investigators had 

 confused "parts of at least two and possibly three quite difi^erent aggre- 

 gations of structures, namely, the pellicle, the trichocysts and the periph- 

 eral portion of the neuromotor system. In addition" they had failed 



